Skolithos

Skolithos (formerly spelled Scolithus or Skolithus[1]) is a common trace fossil ichnogenus that is, or was originally, an approximately vertical cylindrical burrow. It is produced by a variety of organisms in shallow marine environments globally and appear as lineated features in sedimentary rocks.[2]

Skolithos
Temporal range: Early Cambrian–Present
Skolithos trace fossils. Scale bar is 10 mm.
Trace fossil classification
Ichnogenus: †Skolithos
Skolithos from Kraków am See, Germany.

Depositional environments

Skolithos burrows range in age from early Cambrian[2] to the present[3][4] and are found throughout the world. They occur in sediments and sedimentary rocks, primarily sands and sandstones. They are typically marine in origin,[5] and are commonly associated with high-energy environments close to the shoreline.[2] They have also been reported from freshwater lacustrine settings,[6] but have rarely been described from carbonate rocks.[7] Vertical Skolithos burrows can also occur in alluvial sediments such as braided river deposits, where the periodic fluctuation of water is an important factor in the formation of this structure.[8]  This periodic water fluctuation corresponds to tidal activity in shallow marine environments, but also occurs over longer time intervals in alluvial deposits.[8]

Classification and history

The Skolithos linearis ichnogenus was first described in 1840, by Samuel Stehman Haldeman, a renowned Pennsylvania naturalist in the early 19th century, who labeled the structure as the “oldest fossil in the state”.[9] He named the trace fossil Skolithos meaning “worm-stone”, suggesting that it may have been a product of animal activity rather than a floral origin.[9] James Hall published the first illustrations of Haldeman’s discovery in his journal Paleontology of New York Volume I (1847) coining it the Latin term Skolithus.[9]  1943 marked the revival of Haldeman’s research as Benjamin Howell reported the occurrence of the trace fossil in the Hardyston Formation in Pennsylvania.[9] Howell retitled the ichnogenus Skolithos in accordance with the appropriate ichnotaxon nomenclature. In the 1960s structural geologists discovered the use of the trace fossil as a strain marker by which it could record rotation and strain in highly deformed rocks.[9] This led to a series of experiments that extend to present day analyses to determine the extent of the strain marking properties of Skolithos burrows. Skolithos linearis, found in the Blue Ridge Mountain region, is the oldest known trace fossil in Virginia.[10] Trypanites is similar in form but is excavated in hard substrates. Also somewhat similar are Ophiomorpha and Diplocraterion.

Structure and use as a strain marker

Skolithos structure

The structure of the trace fossil is cylindrical and elongated in shape, usually at a perpendicular angle to the surface where it has been deposited. They can reach lengths of up to about 35 cm (14 in)[6] and diameters of up to about 5 cm (2.0 in).[6] The vertical burrows are composed of the same mineralogy as its surrounding matrix which allow it to deform homogenously with the parent rock. Variations in observed Skolithos structures include burrow curvature, angle to the plane of deposition, and size of the fossil’s aperture.[11] Funnel-shaped apertures of Skolithos reflect the filter- and suspension-feeding habits of burrowing genera. The high intensity of bioturbation of these organisms indicate the shallow water paleoenvironment in which the Skolithos burrows formed shortly after the deposition of the bed.[11]

Using Skolithos to evaluate strain

Unstrained Skolithos structures are normal to the plane of the bed.[11] In zones where tectonic deformation is intense, such as thrust zones, the deformed Skolithos burrow can be used to evaluate the local strain on the region.[11] This technique is performed by comparing the angle between the specimen and the bedding surface, with the original 90o geometric relationship. Since the trace fossil shares similar material properties to the surrounding matrix, they are inferred to deform by the same mechanism.[11] This technique can be applied in areas where other strain markers may have been destroyed by tectonic activity or cataclastic flow.[11]

Unit strain ɛ can also be defined using the elongation of the structure:

where

•     ɛ is the unit strain due to elongation

•      l  is the deformed length of the structure

•     lo is the initial length of the structure

The structure length and orientation may be influenced by the directional behaviour of the burrowing organism, therefore observing the widths of the burrow may provide a more precise strain estimation.[12]

Example of strain analysis using Skolithos

The famous "Pipe Rock" of northwest Scotland is a well-known example of Skolithos. The 'pipes' that give the rock its name are closely packed straight Skolithos tubes that were presumably made by a worm-like organism.[13] The Pipe Rock can be found in the Stack of Glencoul region beneath the Moine Thrust Belt, Scotland.[14]  This area which has a history of thrust faulting activity is a highly deformed mylonite zone with a quartzite protolith where many structural geologists have used microstructures such as the Skolithos borings in conjunction with other strain markers, such as quartz vein recrystallization, in order to approximate strain in the region.[14] Using three-dimensional analysis of the strain markers, geologists inferred flattening of the region parallel to the thrust direction, stretching along the vertical strain direction and shortening perpendicular to the foliation of the lithology.[14] The deformation history of the mylonite belt which is characterized by large translation of thrust faults, can be deduced from the apparent clockwise rotation of these structures. Assuming simple shear, the westward displacement of the 800 m thick Moine Thurst mylonites at Loch Eriboll where the average shear strain determined using the trace fossils is approximately 10, was calculated to be about 8 km.[11]

Criticisms and sources of error

Assumptions regarding the undeformed burrow and its geometric relationship cannot directly be determined, and only estimated.[15] While it is common for Skolithos burrows to form normal to the deposition plane, this is not always true, in which case, the ideal, undeformed state can no longer be used as a reference orientation.[15] Shallow depositional sediments are also susceptible to damage by erosion and tectonic stress forces, which can influence average measurements and geometric orientations.[7] Since the rheological properties between the structure and the host rock are usually very similar, observations of the fossils are conducted with the assumption that they have deformed homogeneously, where the deformation forces are distributed evenly along the entire deformation zone.[12] This is directly contradicted by the presence of folding and varying elongation measurements of the fossil at different locations in the same deformation zone. Deformation mechanisms are difficult to distinguish using this strain marker, as the thinning and flattening of the highly deformed rocks where they are found, cannot necessarily be attributed to pure shear since the planes may have simply rotated near parallel to the shear plane.[15] It is therefore only possible to make accurate strain determinations of the host rock provided the correct assumption of the deformation mechanism and original measurements.

References
  1. Gevers, T.W.; Frakes, L.A.; Edwards, L.N.; Marzolf, J.E. (1971). "Trace Fossils in the Lower Beacon Sediments (Devonian), Darwin Mountains, Southern Victoria Land, Antarctica". Journal of Paleontology. 45 (1): 81–94. JSTOR 1302754.
  2. Desjardins, P. R.; Mángano, M.G.; Buatois, L. A.; Pratt, B. R. (2010). "Skolithos pipe rock and associated ichnofabrics from the southern Rocky Mountains, Canada: colonization trends and environmental controls in an early Cambrian sand-sheet complex". Lethaia. 43 (4): 507. doi:10.1111/j.1502-3931.2009.00214.x.
  3. Pemberton, S.G. and Frey, R.W. 1985. The Glossifungites ichnofacies: modern examples from the Georgia coast, U.S.A. In: Curran, H.A. (ed.), Biogenic structures: their use in interpreting depositional environments. Society of Economic Paleontologists and Mineralogists, Special Publication 35, p. 237-259.
  4. Gingras, M.K., Pemberton, S.G., Saunders, T. and Clifton, H.E. 1999. The ichnology of modern and Pleistocene brackish-water deposits at Willapa Bay, Washington; variability in estuarine settings. Palaios, vol. 14, no. 4, p. 352-374.
  5. Trewin, N.H.; McNamara, K.J. (1995). "Arthropods invade the land: trace fossils and palaeoenvironments of the Tumblagooda Sandstone (? late Silurian) of Kalbarri, Western Australia". Transactions of the Royal Society of Edinburgh: Earth Sciences. 85 (3): 177–210. doi:10.1017/s026359330000359x.
  6. Woolfe, K.J. (1990). "Trace fossils as paleoenvironmental indicators in the Taylor Group (Devonian) of Antarctica". Palaeogeography, Palaeoclimatology, Palaeoecology. 80 (3–4): 301–310. Bibcode:1990PPP....80..301W. doi:10.1016/0031-0182(90)90139-X.
  7. Vinn, O.; Wilson, M.A. (2013). "An event bed with abundant Skolithos burrows from the late Pridoli (Silurian) of Saaremaa (Estonia)". Carnets de Géologie. CG2013_L02: 83–87. doi:10.4267/2042/49316. Retrieved 2013-04-04.
  8. Fitzgerald, P.G.; Barrett, P.J. (January 1986). "Skolithos in a Permian braided river deposit, southern Victoria Land, Antarctica". Palaeogeography, Palaeoclimatology, Palaeoecology. 52 (3–4): 237–247. doi:10.1016/0031-0182(86)90049-0.
  9. Knaust, Dirk; Thomas, Roger D.K.; Curran, H. Allen (October 2018). "Skolithos linearis Haldeman, 1840 at its early Cambrian type locality, Chickies Rock, Pennsylvania: Analysis and designation of a neotype". Earth-Science Reviews. 185: 15–31. doi:10.1016/j.earscirev.2018.05.009.
  10. "Blue Ridge". geology.blogs.wm.edu. Retrieved 2017-02-04.
  11. Wilkinson, P.; Soper, N.J.; Bell, A.M. (October 1975). "Skolithos pipes as strain markers in mylonites". Tectonophysics. 28 (3): 143–157. doi:10.1016/0040-1951(75)90033-5.
  12. Waldron, John W.F (January 1988). "Determination of finite strain in bedding surfaces using sedimentary structures and trace fossils: A comparison of techniques". Journal of Structural Geology. 10 (3): 273–281. doi:10.1016/0191-8141(88)90060-0.
  13. Hallam, A. and Swett, K. Trace fossils from the Lower Cambrian pipe rock of the north-west highlands. Scottish Journal of Geology, vol. 2, p. 101-107.
  14. Law, R. D.; Mainprice, D.; Casey, M.; Lloyd, G. E.; Knipe, R. J.; Cook, B.; Thigpen, J. R. (2010). "Moine Thrust zone mylonites at the Stack of Glencoul: I – microstructures, strain and influence of recrystallization on quartz crystal fabric development". Geological Society, London, Special Publications. 335 (1): 543–577. doi:10.1144/SP335.23. ISSN 0305-8719. S2CID 73567897.
  15. McLeish, Andrew J. (December 1971). "Strain analysis of deformed Pipe Rock in the Moine Thrust zone, northwest Scotland". Tectonophysics. 12 (6): 469–503. doi:10.1016/0040-1951(71)90046-1.
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