Phymaturus

Phymaturus is a genus of iguanian lizards of the family Liolaemidae, a family which was traditionally included in the Iguanidae as a subfamily, but more recently was proposed to warrant distinct family status. Phymaturus is the mid-sized genus of its family, with 48 species altogether known as of 2019;[2] new species are still being discovered however.[3]

Phymaturus
Adult male of Phymaturus verdugo
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Suborder: Iguania
Family: Liolaemidae
Genus: Phymaturus
Gravenhorst, 1838[1]
Diversity
48 species
Synonyms

Centrura Bell, 1843

Species of the genus Phymaturus are found in the Andes region south to Patagonia and inhabit a variety of habitats. Their habits are mostly conserved from the ancestral iguanians, in that Phymaturus are generally inhabitants of rocky ground, feed on plants, and give birth to fully developed young.[3]

Systematics

The genus can be divided into two lineages, which probably represent clades:

palluma group

Superciliar scales not imbricate, more than four subocular scales, 3-4 rows of lorilabial scales, mental scale narrower than rostral scale and usually touching the sublabial scales. Tail spines well-developed, two annuli per segment.[3]
  • Phymaturus aguedae Troncoso-Palacios & Esquerré, 2014
  • Phymaturus antofagastensis Pereyra, 1985
  • Phymaturus damasense Troncoso-Palacios & Lobo, 2012
  • Phymaturus extrilidus Lobo et al., 2012
  • Phymaturus loboi Troncoso-Palacios, Ferri-Yanez, Laspiur, & Aguilar, 2019
  • Phymaturus mallimaccii Cei, 1980
  • Phymaturus palluma (Molina, 1782) (= P. flagellifer)
  • Phymaturus punae Cei, Etheridge & Videla, 1983
  • Phymaturus roigorum Lobo & Abdala, 2007
  • Phymaturus timi Hibbard, Nenda, & Lobo, 2019
  • Phymaturus verdugo Cei & Videla, 2003
  • Phymaturus vociferator Pincheira-Donoso, 2004 (= P. dorsimaculatus)

patagonicus group

Superciliar scales elongate and overlapping, one usually unfragmented subocular scale, tail smooth, Meckel's groove fused and closed.[3]
  • Phymaturus calcogaster Scolaro & Cei, 2003
  • Phymaturus ceii Scolaro & Ibargüengoytía, 2007
  • Phymaturus delheyi Ávila et al., 2011[4]
  • Phymaturus indistinctus Cei & Castro, 1973
  • Phymaturus nevadoi Cei & Castro, 1973
  • Phymaturus patagonicus Koslowsky, 1898
  • Phymaturus payuniae Cei & Castro, 1973
  • Phymaturus sitesi Avila et al., 2011[4]
  • Phymaturus somuncurensis Cei & Castro, 1973
  • Phymaturus spurcus Barbour 1921
  • Phymaturus tenebrosus Lobo & Quinteros, 2005
  • Phymaturus zapalensis Cei & Castro, 1973

More species which may or may not belong to the aforementioned species groups:[2]

  • Phymaturus aguanegra Lobo, Laspiur & Acosta, 2013
  • Phymaturus alicahuense Núñez et al., 2010
  • Phymaturus bibronii (Guichenot, 1848)
  • Phymaturus cacivioi Lobo & Nenda, 2015
  • Phymaturus camilae Scolaro, Jara & Pincheira-Donoso, 2013
  • Phymaturus castillensis Scolaro & Pincheira-Donoso, 2010
  • Phymaturus curivilcun Scolaro et al., 2016
  • Phymaturus darwini Núñez et al., 2010
  • Phymaturus denotatus Lobo, Nenda & Slodki, 2012
  • Phymaturus desuetus Scolaro & Tappari, 2009
  • Phymaturus dorsimaculatus Lobo & Quinteros, 2005
  • Phymaturus etheridgei Lobo, Abdala & Valdecantos, 2010
  • Phymaturus felix Lobo, Abdala & Valdecantos, 2010
  • Phymaturus fiambala Lobo, Hibbard, Quipildor, & Valdecantos, 2019
  • Phymaturus laurenti Lobo, Abdala & Valdecantos, 2010
  • Phymaturus manuelae Scalaro & Ibargüengoytía, 2008
  • Phymaturus maulense Núñez et al., 2010
  • Phymaturus querque Lobo, Abdala & Valdecantos, 2010
  • Phymaturus rahuensis González-Marín et al., 2016
  • Phymaturus sinervoi Scolaro, De La Cruz & Ibargüengoytía, 2012
  • Phymaturus tromen Lobo & Nenda, 2015
  • Phymaturus videlai Scolaro & Pincheira-Donoso, 2010
  • Phymaturus williamsi Lobo, Laspiur & Acosta, 2013

Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than Phymaturus.

Footnotes
  1. "Phymaturus ". Dahms Tierleben. www.dahmstierleben.de/systematik/Reptilien/Squamata/Iguania/liolaemidae.
  2. Phymaturus at the Reptarium.cz Reptile Database. Accessed 15 September 2017.
  3. Scolaro et al. (2008).
  4. Avila et al. (2011).

References
  • Avila, Luciano Javier; Perez, Cristian Hernan Fulvio; Perez, Daniel Roberto; Morando, Mariana (2011). "Two new mountain lizard species of the Phymaturus genus (Squamata: Iguania) from northwestern Patagonia, Argentina". Zootaxa 2924: 1-21. (in English with Spanish abstract). PDF abstract.
  • Gravenhorst, J.L.C. (1838). "Beiträge zur genaueren Kenntniss einiger Eidechsgattungen ". Nova Acta Academiae Caesareae Leopoldino-Carolinae 18: 712-784 + Plates LIV-LVI. (Phymaturus, new genus, pp. 749–750). (in German).
  • Scolaro, José Alejandro; Ibargüengoytía, Nora Ruth; Pincheira-Donoso, Daniel (2008). "When starvation challenges the tradition of niche conservatism: On a new species of the saxicolous genus Phymaturus from Patagonia Argentina with pseudoarboreal foraging behaviour (Iguania, Liolaemidae)". Zootaxa 1786: 48-60. (in English with Spanish abstract). PDF abstract.
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