Helicostyla

Helicostyla is a genus of small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily Helicostylinae of the family Camaenidae. Helicostyla is the type genus of the subfamily Helicostylinae.[2] [3]

Helicostyla
A shell of Helicostyla camelopardalis
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Heterobranchia
Superorder:	Eupulmonata
Order:	Stylommatophora
Infraorder:	Helicoidei
Superfamily:	Helicoidea
Family:	Camaenidae
Genus:	Helicostyla Férussac, 1821[1]
Type species
Helix mirabilis Férussac, 1821
Synonyms
Bulimus (Eudoxus) Albers, 1850 Eudoxus Albers, 1850 Helicostyla (Cochlodryas) Martens, 1860· accepted, alternate representation Helicostyla (Dryocochlias) Möllendorff, 1898· accepted, alternate representation (basionym) Helicostyla (Eudoxus) Albers, 1850· accepted, alternate representation Helicostyla (Opalliostyla) Pilsbry, 1896 (invalid: unnecessary replacement name)

Distribution

This genus is endemic to the Philippines.

Species

Species within the genus Helicostyla include:[4][5]

• Helicostyla amagaensis de Chavez, Kendrich, Fontanilla, Batomalaque & Chiba, 2015
• Helicostyla annulata (Sowerby, 1841)
• Helicostyla balteata (Sowerby, 1841)
• Helicostyla bembicodes (Pfeiffer, 1851)
• Helicostyla bullala (Broderip, 1841)
• Helicostyla bustoi (Hidalgo, 1887)
• Helicostyla butleri (Pfeiffer, 1842)
• Helicostyla calamianica (Quadras Moellendorff, 1894)
• Helicostyla camelopardalis (Broderip, 1841)
• Helicostyla canonizadoi (Bartsch, 1932)
• Helicostyla chocolatina (Poppe, Tagaro & Sarino, 2015)
• Helicostyla cincinna (Sowerby, 1840)
• Helicostyla concinna (Sowerby I, 1841)
• Helicostyla decora (A. Adams & Reeve, 1850)
• Helicostyla decorata (Férussac, 1821)
• Helicostyla delicata Fulton, 1903
• Helicostyla diana (Broderip, 1841)
• Helicostyla domingoi (Bartsch, 1919)
• Helicostyla effusa (Pfeiffer, 1842)
• Helicostyla evanescens (Broderip, 1841)
• Helicostyla fastidiosa (Bartsch, 1932)
• Helicostyla fauna (Broderip, 1841)
• Helicostyla gigas (Hidalgo, 1901)
• Helicostyla gilva (Pfeiffer, 1845)
• Helicostyla glaucophthalma (L. Pfeiffer, 1851)
• Helicostyla grandis (Pfeiffer, 1845)
• Helicostyla halichlora (Semper, 1866)
• Helicostyla hololeuca (L. Pfeiffer, 1855)
• Helicostyla hydrophana Sowerby I, 1841
• Helicostyla imperator (Pfeiffer, 1848)
• Helicostyla lacerata (Semper, 1877) (nomen nudum)
• Helicostyla mateoi (Bartsch, 1932)
• Helicostyla mearnsi Bartsch, 1905
• Helicostyla nimbosa (Broderip, 1841)
• Helicostyla nobilis (Reeve, 1848)
• Helicostyla okadai Kuroda, 1932
• Helicostyla olanivanensis (Bartsch, 1913)
• Helicostyla opalina (Sowerby, 1841)
• Helicostyla orbitula (Sowerby, 1841)
• Helicostyla persimilis (Deshayes, 1850)
• Helicostyla phaeostyla (Pfeiffer, 1856)
• Helicostyla pithogaster (Ferussac, 1821)
• Helicostyla polita Reeve, 1848
• Helicostyla pudibunda Semper, 1891
• Helicostyla rehbeini (L. Pfeiffer, 1852)
• Helicostyla rustica (Mousson, 1849)
• Helicostyla saranganica (Hidalgo, 1887)
• Helicostyla smaragdina (Reeve, 1842)
• Helicostyla submirabilis Möllendorff, 1897
• Helicostyla thomsoni (L. Pfeiffer, 1871)
• Helicostyla turbinoides (Broderip, 1841)
• Helicostyla turbo (Pfeiffer, 1845)
• Helicostyla velata (Broderip, 1841)
• Helicostyla weberi (Bartsch, 1919)
• Helicostyla worcesteri Bartsch, 1909

Species brought into synonymy

• Helicostyla aegle (Broderip, 1841): synonym of Cochlostyla aegle (Broderip, 1841)
• Helicostyla bicolorata (Lea, 1840): synonym of Cochlostyla bicolorata (I. Lea, 1840) (superseded combination)
• Helicostyla carinata (Lea, 1840): synonym of Hypselostyla carinata (I. Lea, 1840) (superseded combination)
• Helicostyla cincinniformis (Sowerby, 1841): synonym of Hypselostyla cincinniformis (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla collodes (Sowerby I, 1841): synonym of Cochlostyla collodes (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla curta (Sowerby I, 1841) : synonym of Cochlostyla curta (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla dactylus (Broderip, 1841): synonym of Hypselostyla dactylus (Broderip, 1841) (superseded combination)
• Helicostyla daphnis (Broderip, 1841): synonym of Cochlostyla daphnis (Broderip, 1841) (superseded combination)
• Helicostyla denticulata (Jay, 1839): synonym of Calocochlea denticulata (Jay, 1839) (superseded combination)
• Helicostyla dimera (Jonas, 1846): synonym of Cochlostyla dimera (Jonas, 1846) (superseded combination)
• Helicostyla dubiosa (Pfeiffer, 1846): synonym of Cochlostyla dubiosa (L. Pfeiffer, 1846) (superseded combination
• Helicostyla euconica (Bartsch, 1932): synonym of Cochlostyla effusa (L. Pfeiffer, 1843)
• Helicostyla fenestrata (Sowerby I, 1841): synonym of Pachysphaera fenestrata (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla florida (Sowerby I, 1841): synonym of Cochlodryas florida (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla fulgens (Broderip, 1841): synonym of Cochlostyla fulgens (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla fulgetra (Broderip, 1841): synonym of Cochlostyla fulgetra (Broderip, 1841)
• Helicostyla ignobilis Sowerby I, 1841: synonym of Cochlodryas ignobilis (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla iloconensis (Sowerby I, 1841): synonym of Pachysphaera iloconensis (G. B. Sowerby I, 1841) (misspelling of original genus name, Orustia, Mörch, 1852)
• Helicostyla jonasi (Pfeiffer, 1845): synonym of Cochlostyla jonasi (L. Pfeiffer, 1846) (superseded combination)
• Helicostyla juglans (Pfeiffer, 1842): synonym of Cochlostyla juglans (L. Pfeiffer, 1842) (superseded combination)
• Helicostyla leopardus (Pfeiffer, 1845): synonym of Cochlostyla leopardus (L. Pfeiffer, 1845) (superseded combination)
• Helicostyla leucophaea (Sowerby I, 1841): synonym of Cochlostyla leucophaea (G. B. Sowerby I, 1841) (superseded combination)
• Helicostyla lignaria (Pfeiffer, 1842): synonym of Cochlostyla lignaria (L. Pfeiffer, 1842) (suspected synonym)
• Helicostyla macrostoma (Pfeiffer, 1843): synonym of Cochlostyla macrostoma (L. Pfeiffer, 1843) (superseded combination)
• Helicostyla marinduquensis (Hidalgo, 1887): synonym of Cochlostyla marinduquensis Hidalgo, 1887 (superseded combination)
• Helicostyla metaformis (Ferussac, 1821): synonym of Dryocochlias metaformis (Férussac, 1821) (misspelling of original name, Corasia, Albers, 1850)
• Helicostyla mindoroensis W. J. Broderip, 1892 : synonym of Chrysallis mindoroensis (Broderip, 1841) (unaccepted generic combination)
• Helicostyla mirabilis (Ferussac, 1821): synonym of Cochlostyla mirabilis (Férussac, 1821) (superseded combination)
• Helicostyla montana Semper, 1891: synonym of Cochlostyla montana C. Semper, 1877 (superseded combination)
• Helicostyla monticula (Sowerby I, 1841): synonym of Orustia monticula (G. B. Sowerby I, 1841) (misspelling of original genus name, Orustia, Mörch, 1852)
• Helicostyla pictor (Broderip, 1841): synonym of Cochlostyla pictor (Broderip, 1841) (superseded combination)
• Helicostyla polillensis (Pfeiffer, 1861): synonym of Cochlostyla polillensis (L. Pfeiffer, 1861) accepted as Calocochlea polillensis (L. Pfeiffer, 1861)
• Helicostyla rufogaster (Lesson, 1831): synonym of Cochlostyla rufogaster (Lesson, 1832) (superseded combination)
• Helicostyla sarcinosa (Ferussac, 1821): synonym of Helicobulinus sarcinosa (Férussac, 1821)
• Helicostyla satyra (Broderip, 1841): synonym of Cochlostyla satyrus (Broderip, 1841) (superseded combination)
• Helicostyla simplex (Jonas, 1843): synonym of Cochlostyla simplex (Jonas, 1843) (superseded combination)
• Helicostyla solivaga (Reeve, 1849): synonym of Cochlostyla solivaga (Reeve, 1849) (superseded combination)
• Helicostyla sphaerica (Sowerby I, 1841): synonym of Pachysphaera sphaerica (G. B. Sowerby I, 1841)
• Helicostyla ticaonica (Broderip, 1841): synonym of Cochlostyla ticaonica (Broderip, 1841) (superseded combination)
• Helicostyla ventricosa (Bruguière, 1792): synonym of Cochlostyla ventricosa (Bruguière, 1792) (superseded combination)
• Helicostyla viridostriata (Lea, 1840): synonym of Cochlodryas viridostriata (I. Lea, 1840) (superseded combination)
• Helicostyla woodiana (Lea, 1840): synonym of Cochlostyla woodiana (I. Lea, 1840) (superseded combination)

References

1. Férussac A. É. d'A. de (1821). Histoire naturelle générale et particulière des mollusques terrestres et fluviatiles, Tabl. Limaçons, 28.
2. Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
3. MolluscaBase eds. (2021). MolluscaBase. Helicostyla Férussac, 1821. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=993127 on 2021-02-02
4. Bradybaenidae. Malacolos.com, accessed 16 November 2010.
5. Biolib

• Bank, R. A. (2017). Classification of the Recent terrestrial Gastropoda of the World. Last update: July 16th, 2017

External links

• Férussac, A.E.J.P.F. d'Audebard de. (1821-1822). Tableaux systématiques des animaux mollusques classés en familles naturelles, dans lesquels on a établi la concordance de tous les systèmes; suivis d'un Prodrome général pour tous les mollusques ou fluviatiles, vivantes ou fossiles. Paris, 1821 et 1822.
• Pilsbry, H. A. (1896). On the names of certain subgenera of Helicostyla. The Nautilus. 9(9): 108.
• Albers, J. C. (1850). Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet. Berlin: Enslin. 262 pp
• Kennard, A. S. (1942). The Histoire and Prodrome of Férussac. Part III. The divisional names. Proceedings of the Malacological Society of London. 25 (3): 111-118. London
• Pilsbry, H. A. (1903). The use of the generic name Helicostyla. The Nautilus. 17(5): 58
• Albers, J. C.; Martens, E. von. (1860). Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet von Joh. Christ. Albers. Ed. 2. Pp. i-xviii, 1-359. Leipzig: Engelman

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