Cyrtodactylus

Cyrtodactylus (Greek κυρτος kurtos "curved", from κυπτω kuptō "to stoop"; δακτυλος daktulos "finger, toe") is a diverse genus of Asian geckos, commonly known as bent-toed geckos, bow-fingered geckos, and forest geckos. The genus has at least 300 described species as of 2020,[1] which makes it the largest of all gecko genera.

Cyrtodactylus
Marbled bow-fingered gecko (Cyrtodactylus marmoratus)
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Squamata
Family:	Gekkonidae
Subfamily:	Gekkoninae
Genus:	Cyrtodactylus Gray, 1827
Species
Over 300, see text.

Description

Instead of possessing dilated digits like other geckos, members of Cyrtodactylus have slender, curved toes to which the common names for the genus can be attributed.

Taxonomy

The phylogenetic relationships with this genus and the genus Geckoella has not been resolved to date. Based on morphology, Bauer (2002) suggested that it was a subgenus of Cyrtodactylus but a phylogenetic study by Feng et al. (2007), though with inadequate sampling of taxa, restored it to generic status once again.[2][3] All species of Geckoella have been placed here pending a more definitive study.

Phylogeny

The following phylogeny is from Pyron, et al. (2013),[4] and includes 22 Cyrtodactylus species. Cyrtodactylus is a sister group of Hemidactylus.[4]

Cyrtodactylus

Cyrtodactylus oldhami       Cyrtodactylus ayeyarwadyensis         Cyrtodactylus angularis     Cyrtodactylus jarujini           Cyrtodactylus triedrus         Cyrtodactylus marmoratus         Cyrtodactylus irregularis     Cyrtodactylus consobrinus         Cyrtodactylus annulatus       Cyrtodactylus philippinicus     Cyrtodactylus agusanensis                   Cyrtodactylus intermedius       Cyrtodactylus pulchellus           Cyrtodactylus sermowaiensis     Cyrtodactylus loriae         Cyrtodactylus novaeguineae     Cyrtodactylus tuberculatus             Cyrtodactylus klugei     Cyrtodactylus robustus         Cyrtodactylus tripartitus       Cyrtodactylus epiroticus     Cyrtodactylus louisiadensis

Species

The genus Cyrtodactylus contains more than 300 described species. In 2019, six new species of the Cyrtodactylus intermedius complex were identified in the Cardamom Mountains and Associated Highlands region of Southeast Asia.[5]

Species	Authority	Common name	Geographic range
C. aaroni	Günther & Rösler, 2003[6]		New Guinea
C. adleri	Das, 1997		Nicobar Islands, Andaman Sea, India
C. adorus	Shea, Couper, Wilmer & Amey, 2011 [7]		Queensland, Australia
C. aequalis	Bauer, 2003[8]		Mon State, Myanmar
C. agamensis	(Bleeker, 1860)	Agam bow-fingered gecko	Sumatra, Indonesia
C. agusanensis	Taylor, 1915	Mindanao bow-fingered gecko	Mindanao Island, The Philippines
C. albofasciatus	(Boulenger, 1885)	Boulenger's Indian gecko	western Ghats, India
C. amphipetraeus	Chomdej, Suwannapoom, Pawangkhanant, Pradit, Nazarov, Grismer, & Poyarkov, 2020	Tak bent-toed gecko	Tak, Thailand
C. angularis	(M.A. Smith, 1921)	angulated bow-fingered gecko	Eastern Thailand
C. annandalei	Bauer, 2003[8]	Annandale's leaf-toed gecko	Sagaing Division, Myanmar
C. annulatus	Taylor, 1915[9]	Annulated bow-fingered gecko	Mindanao Island, The Philippines
C. arcanus	Oliver, Richards, & Sistrom 2012 [10][11]		Papua New Guinea
C. astrum	Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012 [12][13]	Starry bent-toed gecko	Southern Thailand, Peninsular Malaysia
C. atremus	Kraus & Weijola, 2019		Papua New Guinea
C. aunglini	Murdoch, Grismer, Wood, Thura, Win, Grismer, Trueblood & Quah, 2018[5]	Kyauk Nagar Cave bent-toed gecko	Myanmar
C. auralensis	Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5]	Phnom Aural bent-toed gecko	Phnom Aural, Cambodia
C. aurensis	Grismer, 2005[14]	Aur Island bent-toed gecko	Aur Island, Malaysia
C. auribalteatus	Sumontha, Panitvong & Deein, 2010[15]	Golden-belted bent-toed gecko	Phitsanulok, Thailand
C. australotitiwangsaensis	Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012 [12]	Southern Titiwangsa bent-toed gecko	Peninsular Malaysia
C. ayeyarwadyensis	Bauer, 2003[8]		Rakhine State, Myanmar
C. badenensis	Nguyen, Orlov & Darevsky, 2006[16]		Southern Vietnam
C. baluensis	Mocquard, 1890[17]	Balu bow-fingered gecko	Mount Kinabalu, Borneo
C. bansocensis	Luu, Nguyen, Le, Bonkowski & Ziegler, 2016[18]	Ban Soc bent-toed gecko	Laos
C. batik	Iskandar, Rachmansah & Umilaela, 2011[19]	Batik bent-toed gecko	Indonesia
C. battalensis	Khan, 1993	Reticulate plump-bodied gecko	Pakistan
C. batucolus	L.L. Grismer, Onn, J.L. Grismer, Wood & Belabut, 2008[20]	Besar Island bent-toed gecko	Pulau Besar of the Water Islands Archipelago off the west coast of Melaka, Peninsular Malaysia
C. bayinnyiensis	Grismer, Wood Jr., Thura, Quah, Murdoch, Grismer, Herr, Lin & Kyaw, 2018	Bayin Nyi Cave bent-toed gecko	Myanmar
C. bhupathyi	Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018[21]		Northeast India
C. bichnganae	Tri & Grismer, 2010	Bich Ngan's bent-toed gecko	Vietnam
C. bidoupimontis	Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler, 2012[22]		Vietnam
C. bintangrendah	Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012[12]	Bintang lowland bent-toed gecko	Peninsular Malaysia
C. bintangtinggi	Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012[12]		Peninsular Malaysia
C. biordinis	Brown & Mccoy, 1980[23]	Guadalcanal bow-fingered gecko	Mount Austen, Guadalcanal, Solomon Islands
C. bobrovi	Nguyen et al., 2015[24]	Bobrov's bent-toed gecko	Vietnam
C. bokorensis	Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5]		Bokor Plateau, Cambodia
C. boreoclivus	Oliver, Krey, Mumpuni & Richards, 2011[25]		Papua New Guinea
C. brevidactylus	Bauer, 2002[26]		Myanmar
C. brevipalmatus	(M.A. Smith, 1923)[27]	Kampuchea bow-fingered gecko, short-hand forest gecko	Khao Luang, Nakhon Si Thammarat, Peninsular Thailand
C. buchardi	David, Teynie & Ohler, 2004[28]		Laos
C. bugiamapensis	Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler, 2012[22]		Vietnam
C. calamei	Luu, Bonkowki, Nguyen, Le, Schneider, Ngo & Ziegler, 2016	Calame's bent-toed gecko	central Laos
C. camortensis	Chandramouli, 2020		Kamorta, Nicobar Islands
C. caovansungi	Orlov & Quang Truong, 2007 [29]		Vietnam
C. capreoloides	Rösler, Richards & Günther, 2007		Papua New Guinea
C. cardamomensis	Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5]	Cardamom Mountains bent-toed gecko	Cardamom Mountains, Cambodia
C. cattienensis	Geissler, Nazarov, Orlov, Böhme, Phung, Nguyen & Ziegler, 2009[30]	Cattien bent-toed gecko	Vietnam
C. cavernicolus	Inger & King, 1962	Sarawak bow-fingered gecko, Niah Cave gecko	Sarawak
C. cayuensis	Li, 2007	Cayu bent-toed gecko	China (Cayu County, Xizang)
C. celatus	Kathriner et al., 2014[31]		Indonesia
C. chamba	AgarwaL, Khandekar & Bauer, 2018	Chamba bent-toed gecko	Himachal Pradesh, India
C. chanhomeae	Bauer, Sumontha & Pauwels, 2003[32]		Thailand
C. chaunghanakwaensis	Grismer, Wood Jr., Thura, Quah, Murdoch, Grismer, Herr, Lin & Kyaw, 2018	Chaunghanakwa Hill bent-toed gecko	Myanmar
C. chauquangensis	Quang, Orlov, et al. 2007[33]		Vietnam
C. chrysopylos	Bauer, 2003[8]		Panluang-Pyadakin Wildlife Sanctuary, Shan State, Myanmar
C. chungi	Ostrowski, Le, Ngo, Pham, Phung, Nguyen & Ziegler, 2021[34]	Chung's bent-toed gecko	Binh Thuan province, Vietnam
C. collegalensis	(Beddome, 1870)	forest spotted gecko, Kollegal ground gecko	India
C. condorensis	(M.A. Smith, 1921)	Pulo Condore bow-fingered gecko, Con Dao bow-fingered gecko	Pulo Condore Island, Vietnam
C. consobrinoides	Annandale, 1905		southern Myanmar
C. consobrinus	W. Peters, 1871	Peters's bow-fingered gecko, thin-banded forest gecko	Malaysian peninsula, Borneo
C. cracens	Batuwita & Bahir, 2005		Sri Lanka
C. crustulus	Oliver, Hartman, Turner, Wilde, Austin & Richards, 2020	Manus bent-toed gecko	Papua New Guinea (Manus Island)
C. cryptus	Heidrich, Rösler, Thanh, Böhme & Ziegler, 2007[35]		Vietnam
C. cucdongensis	Schneider & Phung TM, 2014[36]	Cucdong bent-toed gecko	Vietnam
C. cucphuongensis	Ngo & Onn, 2011[37]		Vietnam
C. culaochamensis	Tri, Grismer, Thai & Wood, 2020	Cù Lao Chàm bent–toed gecko	Vietnam
C. dammathetensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Dammathet cave bent-toed gecko	Myanmar
C. darevskii	Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov, 2014	Darevski's bent-toed gecko	Laos
C. darmandvillei	(Weber, 1890)	Darmandville bow-fingered gecko	Flores, Komodo, Pulau Kalao, Sumbawa, Sikka, Lombok in Indonesia
C. dati	Tri, 2013[39]	Dat bent-toed gecko	Vietnam
C. dattanensis	(Khan, 1980)	plump banded gecko, Khan's bow-fingered gecko	nw Pakistan
C. dattkyaikensis	Grismer, Wood, Quah, Grismer, Thura, Oaks & Lin, 2020	Datt Kyaik Hill bent-toed gecko	Myanmar
C. dayangbuntingensis	Quah, Grismer, Wood Jr. & Sah, 2019		peninsular Malaysia
C. deccanensis	(Günther, 1864)	Günther's Indian gecko	Northern Western Ghats of India
C. derongo	Brown & Parker, 1973	Derongo bow-fingered gecko	Papua, New Guinea
C. deveti	(Brongersma, 1948)	Moluccan bow-fingered gecko	Moluccas, Indonesia
C. doisuthep	Kunya et al. 2014[40]	Doi Suthep bent-toed gecko	Thailand
C. dumnuii	Bauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome & Kunya, 2010[41]	Dumnui's bent-toed gecko	Thailand
C. durio	Grismer, Anuar, Quah, Muin, Onn, Grismer & Ahmad, 2010[42]	Durian bent-toed gecko	Malaysia
C. edwardtaylori	Batuwita & Bahir, 2005		Sri Lanka
C. eisenmanae	Tri, 2008[43]	Eisenman's bent-toed gecko	Vietnam
C. elok	Dring, 1979	Malaysia bow-fingered gecko	western Malaysia
C. epiroticus	Kraus, 2008		Papua New Guinea
C. equestris	Oliver, Richards, Mumpuni & Rosler, 2016		Papua New Guinea
C. erythrops	Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome & Kunya, 2009[44]	Red-eyed bent-toed gecko	Thailand
C. evanquahi	Wood, Grismer, Muin, Anuar & Oaks, 2020	Evan Quah's banded bent-toed gecko	Peninsular Malaysia
C. fasciolatus	(Blyth, 1861)[45]	banded bent-toed gecko	Northwest India
C. feae	(Boulenger, 1893)		Myanmar
C. fraenatus	(Günther, 1864)	Sri Lanka bow-fingered gecko	Sri Lanka
C. fumosus	(F. Müller, 1895)[46]		Sulawesi, Indonesia
C. gansi	Bauer, 2003[8]		Min Dat District, Chin State, Myanmar
C. gialaiensis	Luu, Dung, Nguyen, Le & Ziegler, 2017		Vietnam
C. gordongekkoi	Das, 1994 [47]		Lombok, Indonesia
C. grismeri	Tri, 2008	Grismer's bent-toed gecko	Vietnam
C. guakanthanensis	Grismer, Belabut, Quah, Onn & Wood, 2014[48]	Gua Kanthan bent-toed gecko	Malaysia
C. gubaot	Welton, Siler, Linkem, Diesmos & Brown, 2010[49]		the Philippines
C. gubernatoris	(Annandale, 1913)	Sikkimese bent-toed gecko	Darjeeling, India
C. gunungsenyumensis	Grismer et al., 2016[50]	Gunung Senyum bent-toed gecko	Malaysia
C. guwahatiensis	Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018		India
C. halmahericus	Mertens, 1929 [51]		Indonesia
C. hidupselamanya	Grismer, Wood, Anuar, Grismer, Quah, Murdoch, Muin, Davis, Auilar, Klabacka, Cobos, Aowphol & Sites, 2016	Chiku bent-toed gecko	peninsular Malaysia
C. hikidai	Riyanto A, 2012[52]		Bunguran island, Indonesia
C. himalayanus	Duda & Sani, 1977	Himalaya bow-fingered gecko	Jammu, India
C. himalayicus	(Annandale, 1906)	Himalaya bent-toed gecko	India (West Bengal, Darjeeling)
C. hinnamnoensis	Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler, 2016	Hinnamno bent-toed gecko	Laos
C. hitchi	Riyanto, Kurniati & Engilis, 2016		southeastern Sulawesi, Indonesia
C. hontreensis	Tri, Grismer & Grismer, 2008[53]	Hon Tre bent-toed gecko	Vietnam
C. hoskini	Shea, Couper, Wilmer & Amey, 2011[54]		Queensland, Australia
C. houaphanensis	Schneider, Luu, Sitthivong, Teynié, Le, Nguyen & Ziegler, 2020	Houaphan bent–toed gecko	Laos
C. huongsonensis	Luu, Do & Ziegler, 2011[55]		Vietnam
C. huynhi	Ngo & Bauer, 2008[56]	Huynh's bent-toed gecko	Vietnam
C. ingeri	Hikida, 1990	Sabah bow-fingered gecko	Sabah, Borneo
C. interdigitalis	(Ulber, 1992)	skin-toe forest gecko	Thailand, Laos
C. intermedius	Ulber, 1993	intermediate bow-fingered gecko	Cambodia, Vietnam, and Thailand
C. inthanon	Kunya et al., 2015[57]	Doi Inthanon bent-toed gecko	Thailand
C. irianjayaensis	Rösler, 2001		Papua, New Guinea
C. irregularis	(M.A. Smith, 1921)	irregular bow-fingered gecko	Vietnam
C. jaegeri	Luu, Calame, Bonkowski, T. Nguyen & Ziegler, 2014[58]		Laos
C. jaintiaensis	Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018		India
C. jambangan	Welton, Siler, Diesmos & Brown, 2010[59]		the Philippines
C. jarakensis	L.L. Grismer, Onn, J.L. Grismer, Wood & Belabut, 2008[20]	Jarak Island bent-toed gecko	Pulau Jarak off the west coast of Perak, Peninsular Malaysia
C. jarujini	(Ulber, 1992)	Jarujin's forest gecko	Laos and northern Thailand
C. jatnai	Amarasinghe, Riyanto, Mumpuni & Grismer, 2020	Jatna's bent–toed gecko	west Bali, Indonesia
C. jelawangensis	Grismer, Wood, Anuar, Quah et al., 2014[60]	Jelawang bent-toed gecko	Thai-Malay Peninsula
C. jellesmae	Boulenger, 1897	Kabaena bow-fingered gecko	Masarang in Indonesia
C. jeyporensis	(Beddome, 1877)	Jeypore rock gecko	eastern Ghats, India
C. kazirangaensis	Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018		India
C. khammouanensis	Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov, 2014	Khammouane bent-toed gecko	Laos
C. khasiensis	(Jerdon, 1870)	Khasi Hills bent-toed gecko	India and northern Burma
C. khelangensis	Pauwels, Sumontha, Panitvong & Varaguttanonda, 2014[61]	Lampang bent-toed gecko	Thailand
C. kimberleyensis	Aaron & Doughty, 2012[62]	Kimberley bent-toed gecko, East Montalivet Island gecko	Australia
C. kingsadai	Ziegler, Phung, Le & Nguyen, 2013[63]		Vietnam
C. klugei	Kraus, 2008		Papua New Guinea
C. kohrongensis	Grismer, Onn, Oaks, Neang, Sokun, Murdoch, Stuart & Grismer, 2020	Koh Rong Island bent–toed gecko	Cambodia
C. kunyai	Pauwels, Sumontha, Keeratikiat & Phanamphon, 2014[64]	Kunya's bent-toed gecko	Thailand
C. laangensis	Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5]	Phnom Laang bent-toed gecko	Phnom Laang, Cambodia
C. laevigatus	Darevsky, 1964		Komodo Island
C. langkawiensis	Grismer, Wood, et al., 2012[12]	Langkawi Island bent-toed gecko	Langkawi, Malaysia
C. lateralis	Werner, 1896	Sumatra bow-fingered gecko	Sumatra
C. lawderanus	(Stoliczka, 1871)	Lawder's bent-toed gecko	northwestern India
C. leegrismeri	Chan & Norhayati, 2010[65]	Tenggol Island bent-toed gecko	Malaysia
C. lekaguli	Grismer et al., 2012[12]		Southern Thailand
C. lenggongensis	Grismer, Wood, Anuar, Grismer, Quah, Murdoch, Muin, Davis, Auilar, Klabacka, Cobos, Aowphol & Sites, 2016	Lenggong bent-toed gecko	peninsular Malaysia
C. lenya	Mulcahy, Thura & Zug, 2017	Lenya banded bent-toed gecko	Myanmar
C. limajalur	Davis, Bauer, Jackman, Nashriq & Das, 2019	five-banded bent-toed gecko	East Malaysia
C. linnoensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Linno Cave bent-toed gecko	Myanmar
C. linnwayensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Linn-Way bent-toed gecko	Myanmar
C. lomyenensis	Ngo & Pauwels, 2010[66]	Lomyen bent-toed gecko	Laos
C. loriae	(Boulenger, 1897)	Boulenger's bow-fingered gecko	Papua New Guinea
C. louisiadensis	(DeVis, 1892)	ring-tailed gecko	Louisiade Archipelago, Papua New Guinea
C. macrotuberculatus	Grismer & Ahmad, 2008[67]	tuberculate bent-toed gecko, large tubercled bent-toed gecko	Malaysia
C. maelanoi	Grismer, Rujirawan, Termprayoon, Ampai, Yodthong, Wood, Oaks, & Aowphol, 2020	Mae La Noi bent-toed gecko	Mae Hong Son Province, Thailand
C. majulah	Grismer, Wood & Lim 2012		Singapore; Indonesia
C. malayanus	De Rooij, 1915	Borneo bow-fingered gecko	Borneo
C. mamanwa	Welton, Siler, Linkem, Diesmos & Brown, 2010[49][68]		The Philippines
C. mandalayensis	Mahony, 2009[69]		Myanmar
C. manos	Oliver, Karkkainen, Rösler & Richards, 2019	yellow-snouted bent-toed gecko	Papua New Guinea
C. markuscombaii	Darevsky, Helfenberger, Orlov, & Shah, 1998[70]	striped gecko	Nepal
C. marmoratus	Gray, 1831	marbled bow-fingered gecko	Thailand, Malay Peninsula, Indonesia, Papua New Guinea.
C. martini	Tri, 2011[71]	Martin's bent-toed gecko	Vietnam
C. martinstolli	Darevsky, Helfenberger, Orlov, & Shah, 1998[70]	barred gecko	Nepal
C. matsuii	Hikida, 1990	Hikida's bow-fingered gecko, Matsui's bent-toed gecko	Sabah, Borneo
C. mcdonaldi	Shea, Couper, Wilwer & Amey 2011 [72]		Australia
C. medioclivus	Oliver, Richards & Sistrom, 2012 [73]		Papua New Guinea
C. meersi	L. Grismer, Wood, Quah, Murdoch, M. Grismer, Herr, Espinoza, Brown & Lin, 2018	Bago Yoma bent-toed gecko	Myanmar (Yangon)
C. metropolis	Grismer, Wood, Onn, Anuar & Muin, 2014[74]		Malaysia
C. mimikanus	(Boulenger, 1914)	false bow-fingered gecko, Mimika bow-fingered gecko, Mimika bent-toed gecko	Papua New Guinea
C. minor	Oliver & Richards, 2012[75]		Papua New Guinea
C. mombergi	Grismer, Wood, Quah, Thura, Herr & Lin, 2019	Indawgyi bent-toed gecko	Myanmar
C. montanus	Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018		India
C. muangfuangensis	Sitthivong, Luu, Ha, Nguyen, Le & Ziegler, 2019	Muangfuang bent-toed gecko	Laos
C. multiporus	Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov, 2014	Multipored bent-toed gecko	Laos
C. muluensis	Davis, Bauer, Jackman, Nashriq & Das, 2019	Mulu bent-toed gecko	East Malaysia
C. murua	Kraus & Allison, 2006[76]		Papua New Guinea
C. myaleiktaung	Grismer, Wood, Thura, Win, Grismer, Trueblood & Quah, 2018	Mya Leik Taung bent-toed gecko	Myanmar
C. myintkyawthurai	L. Grismer, Wood, Quah, Murdoch, M. Grismer, Herr, Espinoza, Brown & Lin, 2018	Mt. Popa bent-toed gecko	Myanmar (Mandalay region, Bago Yoma Range, Bago Region)
C. nagalandensis	Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018		India
C. naungkayaingensis	Grismer, Wood, Thura, Quah, Murdoch, Grismer, Herr, Lin & Kyaw, 2018		Myanmar
C. nebulosus	(Beddome, 1870)	Clouded Indian gecko	India
C. nepalensis	Schleich & Kästle, 1998[77]	Nepalese rock gecko	Nepal
C. ngoiensis	Schneider, Luu, Sittivong, Teynie, Le, Nguyen & Ziegler, 2020	Ngoi bent–toed gecko	Laos
C. nicobaricus	Chandramouli, 2020		Nicobar Islands
C. nigriocularis	Nguyễn, Orlov & Darevsky, 2007		southern Vietnam
C. novaeguineae	(Schlegel, 1844)	New Guinea bow-fingered gecko	New Guinea
C. nuaulu	Oliver, Edgar, Mumpuni, Iskandar & Lilley, 2009[78]		Indonesia
C. nyinyikyawi	Grismer, Wood, Thura, Win & Quah, 2019	Shwe Settaw bent-toed gecko	Myanmar
C. oldhami	(Theobald, 1876)	Oldham's bow-fingered gecko	Myanmar and Thailand
C. otai	Nguyen et al., 2015[24]	Ota's bent-toed gecko	Vietnam
C. pageli	Schneider et al., 2011[79]		Laos
C. pantiensis	L.L. Grismer, Onn, J.L. Grismer, Wood & Belabut, 2008[20]	Panti Mountain bent-toed gecko	Gunung Panti Forest Reserve, Johor, Peninsular Malaysia
C. papilionoides	Ulber & Grossmann, 1991	butterfly forest gecko	Thailand
C. papuensis	(Brongersma, 1934)	Papua bow-fingered gecko	Papua New Guinea
C. paradoxus	Darevsky & Szczerbak, 1997		Kieng Giang, Phu Quoc, and Hon Thom Islands in Vietnam
C. payacola	Johnson, Quah, Anuar, Muin, Wood, Grismer, Greer, Onn, Ahmad, Bauer & Grismer, 2012 [80]		Malaysia
C. payarhtanensis	Mulcahy, Thura & Zug, 2017	Tenasserim Mountain bent-toed gecko	southern Myanmar
C. peguensis	(Boulenger, 1893)	Thai bow-fingered gecko, Pegu forest gecko	Thailand, western Malaysia, and Myanmar
C. petani	Riyanto, Grismer & Wood, 2015[81]	Farmer's bent-toed gecko	Indonesia
C. pharbaungensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Pharbaung Cave bent-toed gecko	Myanmar
C. phetchaburiensis	Pawels, Sumontha & Bauer, 2016		Thailand
C. philippinicus	Steindachner, 1867	Philippine bow-fingered gecko	northern Philippines
C. phnomchiensis	Nyang, Henson & Stuart, 2020[82]	Prey Lang bent-toed gecko[83]	Prey Lang, Cambodia
C. phongnhakebangensis	Ziegler, Rösler, Herrmann & Vũ, 2002	Phong Nha-Ke Bang bow-fingered gecko	Vietnam
C. phuketensis	Sumontha, Pawels, Kunya, Nitikul, Samphanthamit & Grismer, 2012 [84]	Phuket bent-toed gecko	Phuket, Thailand
C. phumyensis	Ostrowski, Do, Le, Ngo, Pham, Nguyen, Nguyen & Ziegler, 2020	Phu My bent-toed gecko	Vietnam
C. phuocbinhensis	S. Nguyen, Le, Tran, Orlov, Lathrop, MacCulloch, Le, Jin, L. Nguyen, T. Nguyen, Hoang, Che, Murphy & Zang, 2013 [85]		Vietnam
C. phuquocensis	Tri, Grismer & Grismer, 2010[86][87]	Phu Quoc bent–toed gecko	Vietnam
C. pinlaungensis	Grismer, Wood, Quah, Thura, Oaks & Lin, 2019	Pinlaung bent-toed gecko	Myanmar
C. pronarus	Shea, Couper, Wilmer & Amey, 2011[54]		Australia
C. psarops	Harvey et al., 2015[88]	Speckle-faced bent-toed gecko	Indonesia
C. pseudoquadrivirgatus	Rösler, Nguyễn, Vũ, Ngô & Ziegler, 2008		central Vietnam
C. pubisulcus	Inger, 1958	Inger's bow-fingered gecko	Borneo
C. puhuensis	Nguyen, Yang, Thi Le, Nguyen, Orlov, Hoang, Nguyen, Jin, Rao, Hoang, Che, Murphy & Zhang, 2014[89]	Pù Hu bent-toed gecko	Vietnam
C. pulchellus	Gray, 1827	Malayan forest gecko	Penang, Malaysia
C. pyadalinensis	Grismer, Wood, Thura, Win & Quah, 2019		Myanmar
C. pyinyaungensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Pyinyaung bent-toed gecko	Myanmar
C. quadrivirgatus	Taylor, 1962	Taylor's bow-fingered gecko, four-striped forest gecko, marbled bent-toed gecko	Thailand, Malaysia, Singapore, and Indonesia
C. ramboda	Batuwita & Bahir, 2005		Sri Lanka
C. ranongensis	Sumontha, Pauwels, Panitvong, Kunya & Grismer, 2015[90]	Ranong bent-toed gecko	Thailand
C. redimiculus	King, 1962	Palawan bow-fingered gecko	Palawan, Philippines
C. rex	Oliver, Richards, Mumpuni & Rösler, 2016		Papua New Guinea
C. rishivalleyensis	Agarwal, 2016	Rishi Valley geckoella	India
C. robustus	Kraus, 2008		Papua New Guinea
C. roesleri	Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh & Schmitz, 2010 [91]	Roesler's bent-toed gecko	Vietnam & Laos
C. rosichonarieforum	Riyanto, Grismer & Wood, 2015[92]	Rosichonariefi's bent-toed gecko	Indonesia
C. rubidus	(Blyth, 1861)	red bow-fingered gecko	Andaman Islands; Myanmar
C. rufford	Luu et al., 2016[93]	Rufford bent-toed gecko	Laos
C. russelli	Bauer, 2003[8]		Htamanthi Wildlife Sanctuary, Sagaing Division and Indawgyi Wildlife Sanctuary, Rakhine State, Myanmar
C. sadanensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Sadan Cave bent-toed gecko	Myanmar
C. sadansinensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Sadan Sin Cave bent-toed gecko	Myanmar
C. sadleiri	Wells & Wellington, 1984	Sadleir's bow-fingered gecko	Christmas Island
C. saiyok	Panitvong, Sumontha, Tunprasert & Pauwels, 2014[94]	Sai Yok bent-toed gecko	Thailand
C. salomonensis	Rösler, Richards & Günther, 2007[95]		Solomon Islands
C. samroiyot	Pauwels & Sumontha, 2014[96]	Sam Roi Yot bent-toed gecko	Thailand
C. sangi	Pauwels, Nazarov, Bobrov & Poyarkov, 2018		southern Vietnam
C. sanook	Pauwels, Sumontha, Latinne & Grismer, 2013[97]	Sanook bent-toed gecko	Thailand
C. sanpelensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Sanpel Cave bent-toed gecko	Myanmar
C. semenanjungensis	Grismer & Leong, 2005	Peninsular bent-toed gecko	western Malaysia
C. semiadii	Riyanto, Bauer & Yudha, 2014[98]		Indonesia
C. semicinctus	Harvey et al., 2015[88]	Kerinci bent-toed gecko	Indonesia
C. septentrionalis	Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018		India
C. septimontium	Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5]	Bảy Núi bent-toed gecko	Bảy Núi Hills, Vietnam
C. seribuatensis	Youmans & Grismer, 2006	Seribuat bent-toed gecko	western Malaysia
C. sermowaiensis	(de Rooij, 1915)	DeRooij's bow-fingered gecko	New Guinea
C. serratus	Kraus, 2007[99]		Papua New Guinea
C. sharkari	Grismer et al., 2014[60]	Sharkari's bent-toed gecko	Thai-Malay Peninsula
C. shwetaungorum	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Shwetaung bent-toed gecko	Myanmar
C. sinyineensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Sin Yine Cave bent-toed gecko	Myanmar
C. slowinskii	Bauer, 2002		Myanmar
C. soba	Batuwita & Bahir, 2005	Dumbara bent-toed gecko, Knuckles bent-toed gecko	Sri Lanka
C. sommerladi	Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler, 2016	Sommerlad's bent-toed gecko	central Laos
C. soni	Le, Nguyen, Le & Ziegler, 2016	Son's bent-toed gecko	Vietnam
C. sonlaensis	Nguyen, Pham, Ziegler, Ngo & Le, 2017		northwestern Vietnam
C. soudthichaki	Luu, Calame, Nguyen, Bonkowski & Ziegler, 2015[100]	Soudthichak's bent-toed gecko	Laos
C. speciosus	(Beddome, 1870)	forest spotted gecko	India
C. spelaeus	Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov, 2014		Laos
C. spinosus	Linkem, McGuire, Hayden, Setiadi, Bickford & Brown, 2008		Sulawesi, Indonesia
C. srilekhae	Agarwal, 2016	Bangalore geckoella	India
C. stresemanni	Rösler & Glaw, 2008	Stresemann's bent-toed gecko	Malaysia
C. subsolanus	Batuwita & Bahir, 2005	Sinharaja bent-toed gecko	Sri Lanka
C. sumonthai	Bauer, 2002	Sumontha's gecko	Thailand
C. sumuroi	Welton, Siler, Linkem, Diesmos & Brown, 2010[49]		the Philippines
C. surin	Chan-ard & Makchai (2011)[101]		Thailand
C. sworderi	(M.A. Smith, 1925)	Johore bow-fingered gecko, Kota-tinggi forest gecko, Sworder's bent-toed gecko	Malaysia
C. tahuna	Riyanto, Arida & Koch, 2018[102]		Sangir Island, Indonesia
C. takouensis	Van Tri * Gamble, 2010[103]	Takou bent-toed gecko	Vietnam
C. tamaiensis	(M.A. Smith, 1940)[104]	Tamai bent-toed gecko	Myanmar
C. tambora	Riyanto, Mulyadi, McGuire, Kusrini, Febylasmia, Basyir & Kaiser, 2017	Tambora bent-toed gecko	Sumbawa, Indonesia
C. tanahjampea	Riyanto, Hamidy & McGuire, 2018	Tanahjampea bent-toed gecko	Tanahjampea Island, Indonesia
C. tanim	Nielsen & Oliver, 2017		Papua New Guinea
C. taungwineensis	Grismer, Wood, Quah, Grismer, Thura, Oaks & Lin, 2020	Taung Wine Hill bent-toed gecko	Myanmar
C. tautbatorum	Welton, Siler, Diesmos & Brown, 2009[105]		the Philippines
C. taybacensis	Pham, Le, Ngo, Ziegler & Nguyen, 2019	Taybac bent-toed gecko	Vietnam
C. taynguyenensis	S. Nguyen, Le, Tran, Orlov, Lathrop, Macculloch, Le, Jin, L. Nguyen, T. Nguyen, Hoang, Che, Murphy & Zhang 2013[106]		Vietnam
C. tebuensis	Grismer, Anuar, Muin, Quah & Wood, 2013[107]	Tebu Mountain bent-toed gecko	Malaysia
C. teyniei	David, Nguyen, Schneider & Ziegler, 2011[108]		Laos
C. thathomensis	Nazarov, Pauwels, Konstantinov, Chulisov, Orlov & Poyarkov, 2018	Thathom bent-toed gecko	Laos
C. thirakhupti	Pauwels, Bauer, Sumontha & Chanhome, 2004[109]	Thirakhupt's bent-toed gecko	Thailand
C. thochuensis	Ngo & Grismer, 2012[110]	Thochu bent–toed gecko	Tho Chu Island, Vietnam
C. thuongae	Phung, Schingen, Ziegler & T. Nguyen, 2014[111]		Vietnam
C. thylacodactylus	Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5]		Phnom Dalai, Cambodia
C. tibetanus	Boulenger, 1905		Tibet
C. tigroides	Bauer, Sumontha & Pauwels, 2003[32]	Tiger bent-toed gecko	Thailand
C. timur	Muin, Mohamed, Onn, Sumarli, Loredo & Heinz, 2014[60]	Banjaran Timur bent-toed gecko	Thai-Malay Peninsula
C. tiomanensis	Das & Lim, 2000[112]	Tioman Island bent-toed gecko	Malaysia
C. triedrus	(Günther, 1864)	spotted bent-toed gecko, Sri Lanka gecko, spotted bow-fingered gecko, spotted ground gecko	Sri Lanka
C. trilatofasciatus	Grismer & Wood, 2012[113]	Cameron Highlands bent-toed gecko	Malaysia
C. tripartitus	Kraus, 2008		Papua New Guinea
C. tripuraensis	Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018		India
C. tuberculatus	Lucas & Frost, 1900		Queensland, Australia
C. urbanus	Purkayastha, Das, Bohra, Bauer & Agarwal, 2020	urban bent-toed gecko	India (Assam)
C. varadgirii	Agarwal et al., 2016[114]	Giri's Geckoella	Mumbai, India
C. variegatus	Blyth, 1859	variegated bow-fingered gecko, Moulmein forest gecko	Myanmar and Thailand
C. vilaphongi	Schneider, Nguyen, Le, Nophaseud, Bonkowski & Ziegler, 2014		Laos
C. wakeorum	Bauer, 2003[8]		Southern Rakhine State, Myanmar
C. wallacei	Hayden, Brown, Gillespie, Setiadi, Linkem, Iskandar, Umilaela, Bickford, Riyanto, Mumpuni & McGuire,2008		Sulawesi, Indonesia
C. wangkulangkulae	Sumontha et al., 2014[115]	Wangkulangkul's bent-toed gecko	Thailand
C. wayakonei	Nguyen, Kingsada, Rösler, Auer & Ziegler, 2010[116]		Northern Laos & Yunnan, China [117]
C. welpyanensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Wel Pyan Cave bent-toed gecko	Myanmar
C. wetariensis	Dunn, 1927	Wetar bow-fingered gecko	Wetar, Indonesia
C. yakhuna	(Deraniyagala, 1945)	northern Sri Lanka gecko, spotted bow-fingered gecko, blotch bow-fingered gecko, demon gecko	Sri Lanka
C. yangbayensis	Tri & Onn, 2010[118]	Yangbay bent-toed gecko	Vietnam
C. yathepyanensis	Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38]	Yathe Pyan Cave bent-toed gecko	Myanmar
C. yoshii	Hikida, 1990	Yoshi's bow-fingered gecko	Sabah, Borneo
C. ywanganensis	Grismer, Wood, Thura, Quah, Murdoch, M. Grismer, Herr, Espinoza & Lin, 2018		Myanmar (Shan)
C. zebraicus	(Taylor, 1962)		Thailand
C. zhaoermii	Shi & Zhao 2010 [119]		Tibet, China
C. ziegleri	Nazarov, Orlov, Nguyen & Ho, 2008 [120]		Vietnam
C. zugi	Oliver, Tjaturadi, Mumpuni, Krey & Richards, 2008[121]		Melanesia

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Further reading

• Bauer, Aaron M. (2002). "Two New Species of Cyrtodactylus (Squamata: Gekkonidae) from Myanmar". Proceedings of the California Academy of Sciences. 53 (7): 73–86.
• Gray, J.E. (1827). "A Synopsis of the Genera of Saurian Reptiles, in which some new Genera are indicated, and others reviewed by actual Examination". The Philosophical Magazine, or Annals of Chemistry, Mathematics, Astronomy, Natural History, and General Science. 2: 54–58. (Cyrtodactylus, new genus, p. 56).

External links

• Diversification of Bent-toed Geckos (Cyrtodactylus) on Sumatra and west Java