Cheiracanthium

Yellow sac spider
	C. insulanum, adult male
	Cheiracanthium sp., Pateros, Washington
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Chelicerata
Class:	Arachnida
Order:	Araneae
Infraorder:	Araneomorphae
Family:	Cheiracanthiidae
Genus:	Cheiracanthium; C. L. Koch, 1839[1]
Type species
	C. punctorium; (Villers, 1789)
Species
	212, see text
Synonyms[1]
	Chiracanthops Mello-Leitão, 1942[2]; Helebiona Benoit, 1977[3];

Cheiracanthium, commonly called yellow sac spiders, is a genus of araneomorph spiders in the family Cheiracanthiidae, and was first described by Carl Ludwig Koch in 1839.[4] They are usually pale in colour, and have an abdomen that can range from yellow to beige. Both sexes range in size from 5 to 10 millimetres (0.20 to 0.39 in). They are unique among common house spiders because their tarsi do not point either outward, like members of Tegenaria, or inward, like members of Araneus), making them easier to identify. The name is a reference to the backwardly directed process on the cymbium of the male palp.[5] The species epithet is derived from the Greek Ancient Greek: χείρ, romanized: cheir, meaning "hand", and Acanthium, a genus of thorny-stemmed plants.

Schematic male of Cheiracanthium
a) claws
b) tarsus
c) metatarsus
d) tibia
e) patella
f) femur
g) trochanter
h) coxa
i) pedipalp
k) setae
m) prosoma (cephalothorax)
n) opisthosoma (abdomen)
o) spinnerets

Venom

Though they are beneficial predators in agricultural fields, they are also known to be mildly venomous to humans. Painful bites may be incurred from species such as C. punctorium in Europe, C. mildei in Europe and North America, C. inclusum in the Americas, C. lawrencei in South Africa and C. japonicum in Japan.[6] Cheiracanthium venom is purportedly necrotic, and can cause pain, swelling, and lesions in humans,[6] but the necrotic nature and severity of its bite has been disputed.[7] A study of twenty confirmed Cheiracanthium bites in the United States and Australia found that none resulted in necrosis.[7]

Some of these spiders are attracted to the smell of volatiles in gasoline.[8] An unspecified Cheiracanthium species is reportedly attracted to the smell of petroleum and has caused problems by weaving webs inside the canister vent of particular models of Mazda vehicles, resulting in blockages and build-up of pressure that could potentially cause fuel leakage from the fuel tank and an increased risk of fire. Mazda issued a voluntary recall of Mazda 6 models built between 2010 and 2012, to fit them with software which would alert drivers if problems were developing.[9]

Species

Cheiracanthium is primarily an Old World genus, with many species found from northern Europe to Japan, from Southern Africa to India and Australia. The only known species in the New World are C. inclusum and C. mildei. While the former also occurs in Africa and Réunion, the latter is found in the Holarctic region and Argentina. They can also be found in the lower mainland of British Columbia, Canada. The genus is quite diverse in Africa and at least three or four species are known to occur in Egyptian cotton fields alone.[1] As of September 2019 it contains 212 species, found in the Caribbean, South America, Oceania, Europe, Central America, Africa, Asia, North America, and on Saint Helena:[1]

C. abbreviatum Simon, 1878 – France, Denmark
C. aculeatum Simon, 1884 – Africa
C. aden Lotz, 2007 – Yemen
C. adjacens O. Pickard-Cambridge, 1885 – China (Yarkand), Karakorum
C. africanum Lessert, 1921 – Africa, Madagascar, Réunion
C. aizwalense Biswas & Biswas, 2007 – India
C. aladanense Lotz, 2007 – Yemen
C. albidulum (Blackwall, 1859) – Madeira
C. algarvense Wunderlich, 2012 – Portugal, Spain
C. ambrense Lotz, 2014 – Madagascar
C. ampijoroa Lotz, 2014 – Madagascar
C. andranomay Lotz, 2014 – Madagascar
C. angolense Lotz, 2007 – Angola, Zimbabwe, South Africa
C. angulitarse Simon, 1878 – Spain, France (Corsica), Italy, Hungary, Romania
C. anjozorobe Lotz, 2014 – Madagascar
C. annulipes O. Pickard-Cambridge, 1872 – Spain, Egypt, Israel
C. antungense Chen & Huang, 2012 – Taiwan
C. apia Platnick, 1998 – Samoa
C. approximatum O. Pickard-Cambridge, 1885 – China (Yarkand)
C. ashleyi Lotz, 2014 – Madagascar
C. auenati Caporiacco, 1936 – Libya
C. auriculatum Zhang, Zhang & Yu, 2018 – China
C. bantaengi Merian, 1911 – Indonesia (Sulawesi)
C. barbarum (Lucas, 1846) – Algeria
C. boendense Lotz, 2015 – Congo
C. brevicalcaratum L. Koch, 1873 – Indonesia (Lombok), Western Australia
C. brevidens Kroneberg, 1875 – Central Asia
C. brevispinum Song, Feng & Shang, 1982 – China, Korea
C. campestre Lohmander, 1944 – Sweden, Denmark, [[Central Europe]], Romania, Ukraine, Russia (Europe)
C. canariense Wunderlich, 1987 – Canary Is., Turkey, Egypt
C. catindigae Barrion & Litsinger, 1995 – Philippines
C. caudatum (Thorell, 1887) – Myanmar
C. conflexum Simon, 1906 – India
C. conspersum (Thorell, 1891) – India (Nicobar Is.)
C. crucigerum Rainbow, 1920 – Australia (Norfolk Is.)
C. danieli Tikader, 1975 – India
C. daquilium Barrion & Litsinger, 1995 – Indonesia, Philippines
C. debile Simon, 1890 – Chad, Yemen
C. denisi Caporiacco, 1939 – Ethiopia, Congo
C. dippenaarae Lotz, 2007 – South Africa
C. echinulatum Zhang, Zhang & Yu, 2018 – China
C. effossum Herman, 1879 – Central to eastern Europe
C. elegans Thorell, 1875 – Europe, Turkey, Caucasus, Russia (Europe to south Siberia), Kazakhstan, Central Asia
C. equestre O. Pickard-Cambridge, 1874 – Libya, Egypt
C. erraticum (Walckenaer, 1802) – Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China, Korea, Japan
C. escaladae Barrion, Barrion-Dupo & Heong, 2013 – China
C. eutittha Bösenberg & Strand, 1906 – Taiwan, Japan, Korea?
C. excavatum Rainbow, 1920 – Australia (Norfolk Is.)
C. exilipes (Lucas, 1846) – Algeria
C. exquestitum Zhang & Zhu, 1993 – China
C. falcatum Chen, Huang, Chen & Wang, 2006 – Taiwan
C. falcis Lotz, 2015 – Gabon
C. festae Pavesi, 1895 – Israel
C. fibrosum Zhang, Hu & Zhu, 1994 – China
C. filiapophysium Chen & Huang, 2012 – Taiwan
C. fisheri Lotz, 2014 – Madagascar
C. floresense Wunderlich, 2008 – Azores
C. foordi Lotz, 2015 – South Africa
C. foulpointense Lotz, 2014 – Madagascar
C. fujianense Gong, 1983 – China
C. fulvotestaceum Simon, 1878 – France
C. furax L. Koch, 1873 – Samoa
C. furculatum Karsch, 1879 – Cape Verde Is., Africa, Madagascar, Comoros
C. ghanaense Lotz, 2015 – Ghana
C. gobi Schmidt & Barensteiner, 2000 – China
C. gracile L. Koch, 1873 – Australia (Queensland, New South Wales)
C. gratum Kulczyński, 1897 – Germany, Hungary
C. griswoldi Lotz, 2014 – Madagascar
C. halophilum Schmidt & Piepho, 1994 – Cape Verde Is.
C. haroniense Lotz, 2007 – Zimbabwe
C. himalayense Gravely, 1931 – India
C. hypocyrtum Zhang & Zhu, 1993 – China
C. ienisteai Sterghiu, 1985 – Romania, Albania
C. ilicis Morano & Bonal, 2016 – Spain
C. impressum Thorell, 1881 – Australia (Queensland)
C. incertum O. Pickard-Cambridge, 1869 – Sri Lanka
C. inclusum (Hentz, 1847) – North America, Central America, Caribbean, South America. Introduced to Réunion
C. incomptum (Thorell, 1891) – India (Nicobar Is.)
C. indicum O. Pickard-Cambridge, 1874 – India, Sri Lanka
C. inflatum Wang & Zhang, 2013 – China
C. inornatum O. Pickard-Cambridge, 1874 – India
C. insigne O. Pickard-Cambridge, 1874 – India, Sri Lanka, Thailand, Myanmar, Laos, China
C. insulanum (Thorell, 1878) – Myanmar, Laos to China, Taiwan, Indonesia (Moluccas), Philippines
C. insulare L. Koch, 1866 – Samoa
C. insulare (Vinson, 1863) – Madagascar, Réunion
C. isiacum O. Pickard-Cambridge, 1874 – Libya, Egypt
C. itakeum Barrion & Litsinger, 1995 – Philippines
C. jabalpurense Majumder & Tikader, 1991 – India
C. japonicum Bösenberg & Strand, 1906 – China, Korea, Japan
C. jocquei Lotz, 2014 – Comoros, Madagascar
C. joculare Simon, 1910 – São Tomé and Príncipe
C. jorgeense Wunderlich, 2008 – Azores
C. jovium Denis, 1947 – Egypt
C. kabalense Lotz, 2015 – Uganda
C. kakamega Lotz, 2015 – Kenya
C. kakumense Lotz, 2015 – Ivory Coast, Ghana, Congo
C. kashmirense Majumder & Tikader, 1991 – India
C. kazachstanicum Ponomarev, 2007 – Kazakhstan
C. kenyaense Lotz, 2007 – Africa
C. kibonotense Lessert, 1921 – Ethiopia, Congo, Kenya, Tanzania, Uganda
C. klabati Merian, 1911 – Indonesia (Sulawesi)
C. knipperi Lotz, 2011 – Tanzania
C. kupense Lotz, 2007 – Cameroon
C. lanceolatum Chrysanthus, 1967 – New Guinea
C. lascivum Karsch, 1879 – Russia (Sakhalin), China, Korea, Japan
C. leucophaeum Simon, 1897 – Madagascar
C. ligawsolanum Barrion & Litsinger, 1995 – Philippines
C. liplikeum Barrion & Litsinger, 1995 – Philippines
C. liuyangense Xie, Yin, Yan & Kim, 1996 – China
C. lompobattangi Merian, 1911 – Indonesia (Sulawesi)
C. longimanum L. Koch, 1873 – Australia (Queensland), Tonga, Fiji, Vanuatu, New Caledonia
C. longipes (Thorell, 1890) – Indonesia (Sumatra)
C. longtailen Xu, 1993 – China
C. ludovici Lessert, 1921 – Congo, Kenya, Tanzania, Madagascar
C. lukiense Lotz, 2015 – Congo
C. macedonicum Drensky, 1921 – Italy, Slovenia, Slovakia, Hungary, Romania, Macedonia, Bulgaria
C. madagascarense Lotz, 2014 – Comoros, Madagascar
C. mahajanga Lotz, 2014 – Madagascar
C. malkini Lotz, 2007 – Nigeria
C. mangiferae Workman, 1896 – Singapore, Indonesia (Sumatra)
C. maraisi Lotz, 2007 – Namibia, Botswana
C. margaritae Sterghiu, 1985 – Romania
C. marplesi Chrysanthus, 1967 – New Guinea
C. mayombense Lotz, 2015 – Congo
C. melanostomum (Thorell, 1895) – India, Bangladesh, Myanmar
C. mertoni Strand, 1911 – Indonesia (Aru Is.)
C. mildei L. Koch, 1864 – Europe, North AfricaEurope, North Africa, Turkey, Middle East, Caucasus, Russia (Europe) to Central Asia. Introduced to North America, Argentina
C. minahassae Merian, 1911 – Indonesia (Sulawesi)
C. minshullae Lotz, 2007 – Zimbabwe, Botswana, South Africa
C. molle L. Koch, 1875 – Africa, Saudi Arabia
C. mondrainense Main, 1954 – Australia (Western Australia)
C. mongolicum Schenkel, 1963 – Mongolia
C. montanum L. Koch, 1877 – Europe, Turkey, Caucasus, Iran
C. mordax L. Koch, 1866 – China, Taiwan, Australia to Samoa, New Hebrides, Solomon Is., French Polynesia
C. murinum (Thorell, 1895) – India, Myanmar
C. mysorense Majumder & Tikader, 1991 – India, Bangladesh
C. nalsaroverense Patel & Patel, 1973 – India
C. nervosum Simon, 1909 – Australia (Western Australia)
C. nickeli Lotz, 2011 – Mauritania
C. ningmingense Zhang & Yin, 1999 – China
C. occidentale L. Koch, 1882 – Spain (Minorca), Italy
C. olliforme Zhang & Zhu, 1993 – China
C. oncognathum Thorell, 1871 – Europe
C. pallidum Rainbow, 1920 – Australia (Lord Howe Is.)
C. pauriense Majumder & Tikader, 1991 – India
C. pelasgicum (C. L. Koch, 1837) – Southern and eastern Europe, Turkey, Caucasus, Russia (Europe) to Tajikistan
C. pennatum Simon, 1878 – Southern Europe, Romania
C. pennuliferum Simon, 1909 – Australia (Western Australia)
C. pennyi O. Pickard-Cambridge, 1873 – Europe, Turkey, Caucasus, Russia (Europe to south Siberia), Iran, Central Asia, China
C. peregrinum Thorell, 1899 – Ivory Coast, Nigeria, Cameroon
C. pichoni Schenkel, 1963 – China
C. poonaense Majumder & Tikader, 1991 – India
C. potanini Schenkel, 1963 – China
C. punctipedellum Caporiacco, 1949 – Congo, Rwanda, Kenya
C. punctorium (Villers, 1789) (type) – Europe, Turkey, Caucasus, Russia (Europe to south Siberia), Iran, Central Asia
C. punjabense Sadana & Bajaj, 1980 – India
C. ransoni Lotz, 2014 – Madagascar
C. rehobothense Strand, 1915 – Israel
C. rothi Lotz, 2014 – Madagascar
C. rupicola (Thorell, 1897) – Myanmar, China, Indonesia
C. russellsmithi Lotz, 2007 – Ethiopia
C. rwandense Lotz, 2011 – Rwanda
C. saccharanalis Mukhtar, 2015 – Pakistan
C. sakoemicum Roewer, 1938 – New Guinea
C. salsicola Simon, 1932 – France
C. sambii Patel & Reddy, 1991 – India
C. sansibaricum Strand, 1907 – Ivory Coast to Zanzibar
C. saraswatii Tikader, 1962 – India
C. schenkeli Caporiacco, 1949 – Kenya, Rwanda, Zimbabwe, Botswana, South Africa
C. seidlitzi L. Koch, 1864 – Mediterranean to Central Asia
C. seshii Patel & Reddy, 1991 – India
C. shilabira Lotz, 2015 – Congo, Kenya
C. shiluvanense Lotz, 2007 – South Africa
C. sikkimense Majumder & Tikader, 1991 – India, Bangladesh
C. silaceum Rainbow, 1897 – Australia (New South Wales)
C. simaoense Zhang & Yin, 1999 – China
C. simplex Thorell, 1899 – Cameroon, Nigeria
C. siwi El-Hennawy, 2001 – Egypt
C. solidum Zhang, Zhu & Hu, 1993 – China
C. soputani Merian, 1911 – Indonesia (Sulawesi)
C. spectabile (Thorell, 1887) – Myanmar
C. sphaericum Zhang, Zhu & Hu, 1993 – China
C. stratioticum L. Koch, 1873 – New Zealand, Australia (Tasmania)
C. streblowi L. Koch, 1879 – Russia (Middle and south Siberia)
C. striolatum Simon, 1878 – Western Mediterranean
C. taegense Paik, 1990 – China, Korea, Japan
C. tagorei Biswas & Raychaudhuri, 2003 – Bangladesh
C. taiwanicum Chen, Huang, Chen & Wang, 2006 – China, Taiwan
C. tanmoyi Biswas & Roy, 2005 – India
C. tanzanense Lotz, 2015 – Tanzania
C. taprobanense Strand, 1907 – Sri Lanka
C. tenue L. Koch, 1873 – Australia (Queensland)
C. tetragnathoide Caporiacco, 1949 – Kenya
C. torricellianum Strand, 1911 – New Guinea
C. torsivum Chen & Huang, 2012 – Taiwan
C. triviale (Thorell, 1895) – India, Myanmar
C. trivittatum Simon, 1906 – India
C. truncatum (Thorell, 1895) – Myanmar
C. turanicum Kroneberg, 1875 – Uzbekistan, Tajikistan
C. turiae Strand, 1917 – Thailand to Australia (Queensland)
C. uncinatum Paik, 1985 – China, Korea
C. unicum Bösenberg & Strand, 1906 – Korea, Japan, China, Laos
C. vankhedei Marusik & Fomichev, 2016 – Mongolia
C. vansoni Lawrence, 1936 – Southern Africa
C. verdense Lotz, 2011 – Cape Verde Is.
C. virescens (Sundevall, 1833) – Europe, Caucasus, Russia (Europe to Far East), Iran, China
C. vorax O. Pickard-Cambridge, 1874 – India
C. warsai Mukhtar, 2015 – Pakistan
C. wiehlei Chrysanthus, 1967 – New Guinea
C. wilma (Benoit, 1977) – St. Helena
C. zebrinum Savelyeva, 1972 – Russia (south Siberia), Kazakhstan
C. zhejiangense Hu & Song, 1982 – China, Korea

References

Gloor, Daniel; Nentwig, Wolfgang; Blick, Theo; Kropf, Christian (2019). "Gen. Cheiracanthium C. L. Koch, 1839". World Spider Catalog Version 20.0. Natural History Museum Bern. doi:10.24436/2. Retrieved 2019-10-06.
Bonaldo, A. B.; Brescovit, A. D. (1992). "As aranhas do gênero Cheiracanthium C. L. Koch, 1839 na região neotropical (Araneae, Clubionidae)". Revista Brasileira de Entomologia. 36: 732.
Lotz, L. N. (2007). "The genus Cheiracanthium (Araneae: Miturgidae) in the Afrotropical region. 1. Revision of known species". Navorsinge van die Nasionale Museum Bloemfontein. 23: 4.
Koch, C. L. (1839). Die Arachniden. C. H. Zeh'sche Buchhandlung. pp. 125–158.
Ubick, D.; et al. (2005). Spiders of North America: An Identification Manual. American Arachnological Society.
Papini, R (2012). "Documented bites by a yellow sac spider (Cheiracanthium punctorium) in Italy: a case report". Journal of Venomous Animals and Toxins Including Tropical Diseases. 18 (3): 349–354. doi:10.1590/S1678-91992012000300014.
Vetter, RS; Isbister, GK; Bush, SP; Boutin, LJ (June 2006). "Verified bites by yellow sac spiders (genus Cheiracanthium) in the United States and Australia: where is the necrosis?". The American Journal of Tropical Medicine and Hygiene. 74 (6): 1043–8. doi:10.4269/ajtmh.2006.74.1043. PMID 16760517.
"Gas-loving spider prompts Mazda recall in U.S." Reuters. March 4, 2011.
"Spider invasion prompts Mazda software fix". BBC News. 7 April 2014. Retrieved 9 April 2014.

External links

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[NMBE-1] Gloor, Daniel; Nentwig, Wolfgang; Blick, Theo; Kropf, Christian (2019). "Gen. Cheiracanthium C. L. Koch, 1839". World Spider Catalog Version 20.0. Natural History Museum Bern. doi:10.24436/2. Retrieved 2019-10-06.

[Bona1992-2] Bonaldo, A. B.; Brescovit, A. D. (1992). "As aranhas do gênero Cheiracanthium C. L. Koch, 1839 na região neotropical (Araneae, Clubionidae)". Revista Brasileira de Entomologia. 36: 732.

[Lotz2007-3] Lotz, L. N. (2007). "The genus Cheiracanthium (Araneae: Miturgidae) in the Afrotropical region. 1. Revision of known species". Navorsinge van die Nasionale Museum Bloemfontein. 23: 4.

[Koch1839-4] Koch, C. L. (1839). Die Arachniden. C. H. Zeh'sche Buchhandlung. pp. 125–158.

[5] Ubick, D.; et al. (2005). Spiders of North America: An Identification Manual. American Arachnological Society.

[Papini-6] Papini, R (2012). "Documented bites by a yellow sac spider (Cheiracanthium punctorium) in Italy: a case report". Journal of Venomous Animals and Toxins Including Tropical Diseases. 18 (3): 349–354. doi:10.1590/S1678-91992012000300014.

[Vett2006-7] Vetter, RS; Isbister, GK; Bush, SP; Boutin, LJ (June 2006). "Verified bites by yellow sac spiders (genus Cheiracanthium) in the United States and Australia: where is the necrosis?". The American Journal of Tropical Medicine and Hygiene. 74 (6): 1043–8. doi:10.4269/ajtmh.2006.74.1043. PMID 16760517.

[8] "Gas-loving spider prompts Mazda recall in U.S." Reuters. March 4, 2011.

[9] "Spider invasion prompts Mazda software fix". BBC News. 7 April 2014. Retrieved 9 April 2014.

Cheiracanthium

Venom

Species

See also

References

Further reading

External links