Carditidae

Carditidae is a family of marine bivalve clams of the order Carditida, which was long included in the Venerida. They are the type taxon of the superfamily Carditoidea.

Carditidae
Shells of several Carditidae species
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Bivalvia
Subclass: Heterodonta
Order: Carditida
Superfamily: Carditoidea
Family: Carditidae
Férussac, 1822
Subfamilies

See text

Carditidae is a neglected and poorly classified family.[1][2] It has six subfamilies recognised by the World Register of Marine Species, but the WoRMS has refrained from assigning contents to any family due to ambiguity and overlap in their definitions.[3][2] Additionally, several genera have been or are still considered subgenera of other genera by some authors.

Genera

The genera of Carditidae recognised by the World Register of Marine Species as of April 2019 are:

  • Akardita La Perna, Brunetti & Della Bella, 2018
  • Arcturellina Chavan, 1951
  • Bathycardita Iredale, 1924
  • Beguina Röding, 1798
  • Cardiocardita Anton, 1838
  • Cardita Bruguière, 1792
  • Carditamera
  • Carditellopsis Iredale, 1936
  • Cardites Link, 1807
  • Centrocardita Sacco, 1899
  • †Choniocardia Cossmann, 1904
  • †Claibornicardia Stenzel & Krause, 1957
  • Coripia de Gregorio, 1885
  • Cyclocardia
  • Glans Megerle von Mühlfeld, 1811
  • Glyptoactis Stewart, 1930
  • †Goosensia Cossmann, 1885
  • Hamacuna Cotton, 1931
  • Lazariella Sacco, 1899
  • Megacardita Sacco, 1899
  • Milneria Dall, 1881
  • †Miodomeris Chavan, 1936
  • Miodontiscus Dall, 1903
  • †Paraglans Chavan, 1941
  • Pleuromeris
  • Powellina Huber, 2010
  • Pteromeris
  • Purpurocardia Maxwell, 1969
  • Strophocardia Olsson, 1961
  • Thecalia H. Adams & A. Adams, 1857
  • †Tutcheria Cox, 1946
  • †Venericardia
  • †Venericor Stewart, 1930
  • Vimentum Iredale, 1925

Synonyms:

  • Arcturella Chavan, 1941 is a junior synonym of Arcturellina. It is not to be confused with Arcturella Sars, 1879, which is a junior synonym of the isopod genus Astacilla.
  • Azarella is a junior synonym of Beguina.
  • Agaria, Cardiocardites [sic] and Divergidens are junior synonyms of Cardiocardita.
  • Actinobolus Mörch, 1853 is a junior synonym of Cardites. It is not to be confused with the rhinoceros beetle genus Actinobolus Westwood, 1842.
  • Ceropsis Dall, 1871 is a junior synonym of Milneria. It should not be confused with the true weevil genus Ceropsis.
  • Miodon Carpenter, 1863 is a junior synonym of Miodontiscus. It should not be confused with Miodon Duméril, 1859, which is a junior synonym of the mole viper genus Polemon.

Notes:

  • In some classifications, Bathycardita is a subgenus of Cardiocardita.[4]
  • In some classifications, Centrocardita is a subgenus of Glans.[5]
  • In some classifications, Claibornicardia is a subgenus of either Glyptoactis or Venericardia.[6]
  • In some classifications, Vimentum is a subgenus of Cyclocardia.[7]
  • In some classifications prior to the 2000s, Carditella was included in Carditidae,[8] but it has since been moved to Condylocardiidae,[9] along with its former junior synonym/subgenus (now independent genus) Carditellona.[10]
  • Increasingly in several 2010s classifications, the subgenus Venericardia (†Baluchicardia) Rutsch and Schenck 1943[11] is considered an independent genus.[12][13][14]

Subfamilies

Chavan, 1969

The original six-subfamily system was first proposed by Chavan in 1969, contain the following genera (some of which are now synonyms or classified in other families):[12]

Carditamerinae Chavan, 1969

  • Arcturellina
  • Bathycardita
  • Cardiocardita
  • Carditamera
  • Carditella
  • Carditellona
  • Carditellopsis
  • Centrocardita
  • Choniocardia
  • Cossmannella
  • Cretocardia
  • Cyclocardia
  • Fenestricardita
  • Glans
  • Goosensia
  • Izumicardia
  • Lazariella
  • Miodontiscus
  • Pleuromeris
  • Plionema
  • Scalaricardita
  • Tutcheria
  • Vetericardiella
  • Vimentum

Carditesinae Chavan, 1969

Carditinae Férussac, 1822

Miodomeridinae Chavan, 1969

Thecaliinae Dall, 1903

  • Milneria
  • Thecalia

Venericardiinae Chavan, 1969

  • Leuroactis
  • Megacardita
  • Pacificor
  • Trapezicardita
  • Venericardia
  • Venericor

Perez, 2019

In 2019, D. E. Pérez proposed a seven-subfamily system for Carditidae.[15]

Carditamerinae

Carditinae

Miodomeridinae

Palaeocarditinae Chavan, 1969

  • *Palaeocardita
  • *Schizocardita

Scalaricarditinae Pérez, 2019

  • Coripia
  • *Scalaricardita
  • Vimentum

Thecaliinae

  • Powellina
  • Thecalia

Venericardiinae

* = not yet recognised by the World Register of Marine Species.

In addition, Pérez proposed possible affiliations for the genera not included in his subfamilies:[15]

  • Carditidae: Carditomantea?
  • Carditinae: Goosensia?
  • Cyclocardida: Ainicardita, Crassicardia, Lunulicardita
  • Eucarditidae: Pseudocardia? (some species)
  • Miodomeridinae: Miodontiscus
  • Scalaricarditinae: Malarossia, Akardita?, Choniocardia (could be Cyclocardida)
  • Stem-Carditidae: Fenestricardita?, Trapezicardita?, Gujocardita? (could be Permophoridae), Septocardia? (could be Cardiidae)
  • Stem-Eucarditidae or Stem-Carditida: Izumicardia
  • Venericardiini: Cossmannella, Cretocardia, Lazariella
  • Venericardiinae: Cycloglans, Strophocardia, Maghrebella?, Subvenericardia?
  • Uncertain: Milneria, Xenocardita

In some classifications prior to the 2000s, Carditidae used to contain the seventh subfamily Carditellinae, until it was reallocated to Condylocardiidae.[8]

References
  1. Pérez, Damián Eduardo (2019-01-15). "Phylogenetic relationships of the family Carditidae (Bivalvia: Archiheterodonta)". Journal of Systematic Palaeontology. 0 (16): 1139–1175. doi:10.1080/14772019.2018.1532463. ISSN 1477-2019. S2CID 92392950.
  2. Perna, Rafael La; Brunetti, Mauro M.; Bella, Giano Della (2018-02-13). "Systematic position of two Pliocene carditids with description of Akardita n. gen. and A. iberica n. sp. (Bivalvia: Carditidae)". Zootaxa. 4379 (2): 223. doi:10.11646/zootaxa.4379.2.4. ISSN 1175-5334. PMID 29689985.
  3. "WoRMS - World Register of Marine Species - Carditidae Férussac, 1822". www.marinespecies.org. Retrieved 2019-04-11.
  4. "Fossilworks: Cardiocardita (Bathycardita)". fossilworks.org. Retrieved 2019-04-11.
  5. "Fossilworks: Glans (Centrocardita)". fossilworks.org. Retrieved 2019-04-11.
  6. "PBDB". paleobiodb.org. Retrieved 2019-04-11.
  7. "PBDB". www.paleobiodb.org. Retrieved 2019-04-11.
  8. Bernard, F. R.; Cai, Ying-Ya; Morton, Brian (1993-06-01). A Catalogue of the Living Marine Bivalve Molluscs of China. Hong Kong University Press. p. 12. ISBN 9789622093249.
  9. "WoRMS - World Register of Marine Species - Carditella E. A. Smith, 1881". www.marinespecies.org. Retrieved 2019-04-11.
  10. "WoRMS - World Register of Marine Species - Carditellona Iredale, 1936". www.marinespecies.org. Retrieved 2019-04-11.
  11. "PBDB". www.paleobiodb.org. Retrieved 2019-04-11.
  12. Pérez, Damián Eduardo (2019-01-15). "Phylogenetic relationships of the family Carditidae (Bivalvia: Archiheterodonta)". Journal of Systematic Palaeontology. 0 (16): 1139–1175. doi:10.1080/14772019.2018.1532463. ISSN 1477-2019. S2CID 92392950.
  13. Lockwood, Rowan; McClure, Kate J. (May 2015). "Relationships among Venericardia (Bivalvia: Carditidae) on the U.S. Coastal Plain during the Paleogene". Journal of Paleontology. 89 (3): 2. doi:10.1017/jpa.2015.23. ISSN 0022-3360. S2CID 4483535.
  14. Pérez, Damián E.; del Río, Claudia J. (November 2017). "The family Carditidae (Bivalvia) in the early Danian of Patagonia (Argentina)". Journal of Paleontology. 91 (6): 1148–1165. doi:10.1017/jpa.2017.69. ISSN 0022-3360.
  15. Pérez, Damián Eduardo (2019-01-15). "Phylogenetic relationships of the family Carditidae (Bivalvia: Archiheterodonta)". Journal of Systematic Palaeontology. 0 (16): 1139–1175. doi:10.1080/14772019.2018.1532463. ISSN 1477-2019. S2CID 92392950.
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