Bopyridae

The Bopyridae are a family of isopod crustaceans in the suborder Cymothoida. There are 1223 individual species contained in this family.[1][2] Members of the family are ectoparasites of crabs and shrimps. They live in the gill cavities or under the carapace where they cause a noticeable swelling. Fossil crustaceans have occasionally been observed to have a similar characteristic bulge.[3]

Bopyridae
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Malacostraca
Order:	Isopoda
Suborder:	Cymothoida
Infraorder:	Epicaridea
Superfamily:	Bopyroidea
Family:	Bopyridae Rafinesque, 1815

Shrimp displaying bulge indicating bopyrid parasitism

Genera

• Acrobelione Bourdon, 1981
• Albunione Markham & Boyko, 1999
• Allathelges Kazmi & Markham, 1999
• Allobopyrus Bourdon, 1983
• Allodiplophryxus Markham, 1985
• Allokepon Markham, 1982
• Allorbimorphus Bourdon, 1976
• Anacepon Nierstrasz & Brender à Brandis, 1931
• Anathelges Bonnier, 1900
• Anchiarthrus Markham, 1992
• Anisarthrus Giard, 1907
• Anisorbione Bourdon, 1981
• Anomophryxus Shiino, 1937
• Anuropodione Bourdon, 1967
• Apocepon Nierstrasz & Brender à Brandis, 1930
• Apophrixus Nierstrasz & Brender à Brandis, 1931
• Aporobopyrina Shiino, 1934
• Aporobopyroides Nobili, 1906
• Aporobopyrus Nobili, 1906
• Argeia Dana, 1853
• Argeiopsis Kensley, 1974
• Astalione Markham, 1975
• Asymmetrione Codreanu, Codreanu & Pike, 1965
• Asymmetrorbione Boyko, 2003
• Athelges Gerstaeker, 1862
• Atypocepon Nierstrasz & Brender à Brandis, 1931
• Azygopleon Markham, 1985
• Balanopleon Markham, 1974
• Bathione Román-Contreras & Boyko, 2007
• Bathygyge Hansen, 1897
• Bopyrella Bonnier, 1900
• Bopyrina Kossmann, 1881
• Bopyrinella Nierstrasz & Brender à Brandis, 1925
• Bopyrinina Shiino, 1933
• Bopyrione Bourdon & Markham, 1980
• Bopyrissa Nierstrasz & Brender à Brandis, 1931
• Bopyroides Stimpson, 1864
• Bopyrophryxus Codreanu, 1965
• Bopyrosa Nierstrasz & Brender à Brandis, 1923
• Bopyrus Latreille, 1802
• Cancricepon Giard & Bonnier, 1887
• Carcinione Bourdon, 1983
• Cardiocepon Nobili, 1906
• Castrione Lima, 1980
• Cataphryxus Shiino, 1936
• Coxalione Bourdon, 1977
• Cryptobopyrus Schultz, 1977
• Dactylokepon Stebbing, 1910
• Dicropleon Markham, 1972
• Diplophryxus Richardson, 1904
• Discomorphus Markham, 2008
• Discorsobopyrus Boyko, 2004
• Entophilus Richardson, 1903
• Eophrixus Caroli, 1930
• Epicepon Nierstrasz & Brender à Brandis, 1931
• Epipenaeon Nobili, 1906
• Epiphrixus Nierstraz & Brender à Brandis, 1932
• Eragia Markham, 1994
• Ergyne Risso, 1816
• Eriphrixus Markham, 1990
• Falsanathelges Boyko & WIlliams, 2003
• Filophryxus Bruce, 1972
• Galathocrypta Román-Contreras & Soto, 2002
• Gareia Bourdon & Bruce, 1983
• Gigantione Kossmann, 1881
• Goleathopseudione Román-Contreras, 2008
• Grapsicepon Giard & Bonnier, 1887
• Gyge Cornalia & Panceri, 1861
• Hemiarthrus Giard & Bonnier, 1887
• Hemicepon Lemos de Castro & Brasil-Lima, 1980
• Hemiphryxus Bruce, 1978
• Heterocepon Shiino, 1936
• Hypercepon Danforth, 1972
• Hyperphrixus Nierstrasz & Brender à Brandis, 1931
• Hypocepon Nierstrasz & Brender à Brandis, 1930
• Hypohyperphrixus Nierstrasz & Brender à Brandis, 1932
• Hypophryxus Shiino, 1934
• Ionella Bonnier, 1900
• Kepon Duvernoy, 1840
• Kolourione Markham, 1978
• Leidya Cornalia & Panceri, 1861
• Litobopyrus Markham, 1982
• Lobocepon Nobili, 1905
• Loki Markham, 1972
• Mediophrixus Markham, 1990
• Megacepon George, 1947
• Mesocepon Shiino, 1951
• Mesophryxus Bruce, 1973
• Metacepon Nierstrasz & Brender à Brandis, 1931
• Metaphrixus Nierstrasz & Brender à Brandis, 1931
• Metathelges Nierstrasz & Brender à Brandis, 1923
• Minicopenaeon Bourdon, 1981
• Miophrixus Barnard, 1955
• Munidion Hansen, 1897
• Neophryxus Bruce, 2007
• Nikione Kensley, 1974
• Ogyridione Markham, 1988
• Onkokepon An, Yu & Li, 2006
• Onychocepon Pérez, 1921
• Orbimorphus Richardson, 1910
• Orbione Bonnier, 1900
• Orophryxus Bruce, 1972
• Orthione Markham, 1988
• Ovobopyrus Markham, 1985
• Ovoionella Boyko, 2004
• Pagurion Shiino, 1933
• Pagurocryptella Boyko & Williams, 2010
• Palaemonellione Markham, 1989
• Parabopyrella Markham, 1985
• Parabopyriscus Markham, 1985
• Parabopyrus Shiino, 1934
• Paracepon Nierstrasz & Brender à Brandis, 1931
• Paragigantione Barnard, 1920
• Parapagurion Shiino, 1933
• Parapenaeon Richardson, 1904
• Parapenaeonella Shiino, 1949
• Parapleurocrypta Chopra, 1923
• Parapleurocryptella Bourdon, 1972
• Parapseudione Nierstrasz & Brender à Brandis, 1931
• Parargeia Hansen, 1897
• Parasymmetrione An, Markham & Yu, 2010
• Parasymmetrorbione An, Boyko & Li, 2013[4]
• Parathelges Bonnier, 1900
• Parione Richardson, 1910
• Parionella Nierstrasz & Brender à Brandis, 1923
• Parionina Nierstrasz & Brender à Brandis, 1929
• Parioninella Nierstrasz & Brender à Brandis, 1930
• Pauperella Nierstrasz & Brender à Brandis, 1929
• Phyllodurus Stimpson, 1857
• Pleurocrypta Hesse, 1865
• Pleurocryptella Bonnier, 1900
• Pleurocryptina Nierstrasz & Brender à Brandis, 1929
• Pliophrixus Caroli, 1930
• Pontobopyrus Markham, 1979
• Probopyria Markham, 1985
• Probopyrinella Nierstrasz & Brender à Brandis, 1929
• Probopyrione Bourdon, 1983
• Probopyriscus Markham, 1982
• Probopyrus Giard & Bonnier, 1888
• Probynia Bourdon & Bruce, 1983
• Procepon Shiino, 1937
• Progebiophilus Codreanu & Codreanu, 1963
• Propseudione Shiino, 1933
• Pseudione Kossmann, 1881
• Pseudionella Shiino, 1949
• Pseudostegias Shiino, 1933
• Rhopalione Pérez, 1920
• Robinione Boyko, Williams & Shields, 2017[5]
• Schizobopyrina Markham, 1985
• Scyracepon Tattersall, 1905
• Shiinoella Bourdon, 1972
• Stegias Richardson, 1904
• Stegoalpheon Chopra, 1923
• Synsynella Hay, 1917
• Trapezicepon Bonnier, 1900
• Tylokepon Stebbing, 1904
• Upogebione Markham, 1985
• Upogebiophilus Nobili, 1906
• Urobopyrus Richardson, 1904

References

1. Chris Boyko & Marilyn Schotte (2011). M. Schotte; C. B. Boyko; N. L. Bruce; G. C. B. Poore; S. Taiti & G. D. F. Wilson (eds.). "Bopyridae". World Marine, Freshwater and Terrestrial Isopod Crustaceans database. World Register of Marine Species. Retrieved June 12, 2011.
2. "WoRMS taxon details, Bopyridae Rafinesque, 1815". World Register of Marine Species. 2019-06-24.
3. Virginia Institute of Marine Science
4. An, J., Bokyo, C. & Li, X. (2013). A new genus and species of parasitic isopod (Isopoda: Bopyridae: Orbioninae) from Chinese waters, with a new locality and host record for Orbione halipori Nierstrasz & Brender à Brandis, 1923. Systematic Parasitology, 84(2), 149–156.
5. Boyko, C. B., Williams, J. D. & Shields, J. D. (2017). Parasites (Isopoda: Epicaridea and Nematoda) from Ghost and Mud Shrimp (Decapoda: Axiidea and Gebiidea) with Descriptions of a New Genus and a New Species of Bopyrid Isopod and Clarification of Pseudione Kossmann, 1881. Zootaxa, 4365, 251–301.