Barnadesioideae

Barnadesioideae
	Chuquiraga oppositifolia
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Asterids
Order:	Asterales
Family:	Asteraceae
Subfamily:	Barnadesioideae; K.Bremer & R.K.Jansen
Tribe:	Barnadesieae; D.Don
Genera
	9, see text

Barnadesioideae is a subfamily of flowering plants in the aster family, Asteraceae. It comprises a single tribe, the Barnadesieae. The subfamily is endemic to South America. Molecular evidence suggests it is a basal clade within the family,[1][2] and it is monophyletic.[3]

The subfamily includes species of annual and perennial herbs, shrubs, and trees up to 30 meters tall. Most are likely pollinated by hummingbirds.[3]

Phylogeny

Genetic analysis has included increasing numbers of taxa and involved a growing number of genes. The relationships of the genus Schlechtendalia could so far not be resolved. The following tree reflects the current scientific insights and Schlechtendalia can be found in two possible positions.[3]

Calyceraceae

Asteraceae

subfamily Barnadesioideae

? Schlechtendalia luzulifolia ?

? Schlechtendalia luzulifolia ?

	Huarpea andina

	Barnadesia

Dasyphyllum subgenus Dasyphyllum

Dasyphyllum subgenus Archidasyphyllum

	Fulcaldea laurifolia

	Arnaldoa

Duseniella patagonica

	Doniophyton

	Chuquiraga

subfamily Famatinanthoideae

	subfamily Mutisioideae

	all other Asterid subfamilies

References

Urtubey, E. and T. F. Stuessy. (2001). New hypotheses of phylogenetic relationships in Barnadesioideae (Asteraceae) based on morphology. Taxon 50(4) 1043–66.
Timme, R. E., et al. (2007). A comparative analysis of the Lactuca and Helianthus (Asteraceae) plastid genomes: identification of divergent regions and categorization of shared repeats. American Journal of Botany 94(3) 302–12.
Gruenstaeudl, Michael; Urtubey, Estrella; Jansen, Robert K.; Samuel, Rosabelle; Barfuss, Michael H.J.; Stuessy, Tod F. (2009). "Phylogeny of Barnadesioideae (Asteraceae) inferred from DNA sequence data and morphology". Molecular Phylogenetics and Evolution. 51 (3): 572–87. doi:10.1016/j.ympev.2009.01.023. PMID 19264147.

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[1] Urtubey, E. and T. F. Stuessy. (2001). New hypotheses of phylogenetic relationships in Barnadesioideae (Asteraceae) based on morphology. Taxon 50(4) 1043–66.

[2] Timme, R. E., et al. (2007). A comparative analysis of the Lactuca and Helianthus (Asteraceae) plastid genomes: identification of divergent regions and categorization of shared repeats. American Journal of Botany 94(3) 302–12.

[gru-3] Gruenstaeudl, Michael; Urtubey, Estrella; Jansen, Robert K.; Samuel, Rosabelle; Barfuss, Michael H.J.; Stuessy, Tod F. (2009). "Phylogeny of Barnadesioideae (Asteraceae) inferred from DNA sequence data and morphology". Molecular Phylogenetics and Evolution. 51 (3): 572–87. doi:10.1016/j.ympev.2009.01.023. PMID 19264147.