Archisargidae

Archisargidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods. It is part of the extinct superfamily Archisargoidea. Most members of the family are known from the Callovian-Oxfordian Daohugou biota of Inner Mongolia, China, and the equivalently aged Karabastau Formation of Kazakhstan. The family has been found to be paraphyletic with respect to Eremochaetidae in a cladistic analysis.[1]

Archisargidae
Temporal range: Callovian–Aptian
Archirhagio mostovskii (top) and A. zhangi (bottom)
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Superfamily: †Archisargoidea
Family: †Archisargidae
Rohdendorf 1962
Genera

See text

Male Mesosolva sp.

Taxonomy
  • †subfamily Archisarginae Rohdendorf 1962
    • †Archirhagio Rohdendorf 1938[2]
    • †Archisargus Rohdendorf 1938
      • †Archisargus maximus Mostovski 1997[6] Karabastau Formation, Kazakhstan, Oxfordian
      • †Archisargus pulcher Rohdendorf 1938 Karabastau Formation, Kazakhstan, Oxfordian
      • †Archisargus spurivenius Zhang et al. 2007a[7] Daohugou, China, Callovian
      • †Archisargus strigatus Zhang et al. 2007a[7] Daohugou, China, Callovian
    • †Calosargus Mostovski 1997[6]
      • †Calosargus (Calosargus) Mostovski 1997
        • †Calosargus (Calosargus) antiquus Zhang et al. 2007b[8] Daohugou, China, Callovian
        • †Calosargus (Calosargus) bellus Zhang et al. 2007b[8] Daohugou, China, Callovian
        • †Calosargus (Calosargus) daohugouensis Zhang et al. 2007b[8] Daohugou, China, Callovian
        • †Calosargus (Calosargus) hani Zhang et al. 2007b[8] Daohugou, China, Callovian
        • †Calosargus (Calosargus) niger Mostovski 1997[6] Karabastau Formation, Kazakhstan, Oxfordian
        • †Calosargus (Calosargus) talbragarensis Oberprieler and Yeates 2012[9] Talbragar Fossil Bed, Australia, Tithonian
        • †Calosargus (Calosargus) tatianae Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian
        • †Calosargus (Calosargus) tenuicellulatus Zhang et al. 2007b Daohugou, China, Callovian
        • †Calosargus (Calosargus) validus Zhang et al. 2007b Daohugou, China, Callovian
      • †Calosargus (Pterosargus) Mostovski 1997
        • †Calosargus (Pterosargus) sinicus Zhang 2010[10] Daohugou, China, Callovian
        • †Calosargus (Pterosargus) thanasymus Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian
      • †Calosargus tatianae Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian
    • †Flagellisargus Zhang 2012[11]
      • †Flagellisargus robustus Zhang 2012 Daohugou, China, Callovian
      • †Flagellisargus sinicus Zhang 2012 Daohugou, China, Callovian
      • †Flagellisargus venustus Zhang 2012 Daohugou, China, Callovian
    • †Mesosolva Hong 1983[12]
      • †Mesosolva angustocellulata Mostovski 1996[13] Karabastau Formation, Kazakhstan, Oxfordian
      • †Mesosolva balyshevae Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
      • †Mesosolva daohugouensis Zhang and Zhang 2003[4] Daohugou, China, Callovian
      • †Mesosolva dolosa Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
      • †Mesosolva hennigi Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
      • †Mesosolva huabeiensis Hong 1983 Haifanggou Formation, China, Callovian
      • †Mesosolva imperfecta Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
      • †Mesosolva karataviensis Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
      • †Mesosolva longivena Mostovski 1996 Shar Teeg, Mongolia, Tithonian
      • †Mesosolva parva Hong 1983 Haifanggou Formation, China, Callovian
      • †Mesosolva rohdendorfi Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
      • †Mesosolva sinensis Zhang et al. 2010[14] Daohugou, China, Callovian
      • †Mesosolva zhangae Zhang 2012[15] Daohugou, China, Callovian
    • †Novisargus Zhang 2014[5]
      • †Novisargus rarus Zhang 2014 Daohugou, China, Callovian
    • †Origoasilus Zhang et al. 2011[16]
    • †Parvisargus Mostovski 1996[17]
      • †Parvisargus malus Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
      • †Parvisargus peior Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
    • †Sharasargus Mostovski 1996[17]
      • †Sharasargus fortis Zhang et al. 2008[18] Daohugou, China, Callovian
      • †Sharasargus maculus Zhang 2014[5] Daohugou, China, Callovian
      • †Sharasargus oresbius Ren 1998[19] Yixian Formation, China, Aptian
      • †Sharasargus ruptus Mostovski 1996 Shar Teeg, Mongolia, Tithonian
      • †Sharasargus spiniger Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian
    • †Sinallomyia Zhang 2012[15]
      • †Sinallomyia ruderalis Ren 1998[19] Yixian Formation, China, Aptian
    • †Tabanisargus Zhang 2014[5]
      • †Tabanisargus daohugouensis Zhang 2014 Daohugou, China, Callovian
  • †subfamily Uranorhagioninae Zhang et al. 2010[20] Grimaldi 2016 suggests that this grouping does not belong to the superfamily.[1]
    • †Daohugosargus Zhang 2012[15]
      • †Daohugosargus eximius (Zhang et al. 2008)[18] Daohugou, China, Callovian
    • †Uranorhagio Zhang et al. 2010[20]
      • †Uranorhagio asymmetricus Zhang et al. 2010 Daohugou, China, Callovian
      • †Uranorhagio daohugouensis Zhang et al. 2010 Daohugou, China, Callovian
      • †Uranorhagio deviatus Zhang et al. 2010 Daohugou, China, Callovian

References
  1. Grimaldi, David A.; Barden, Phillip (2016-09-29). "The Mesozoic Family Eremochaetidae (Diptera: Brachycera) in Burmese Amber and Relationships of Archisargoidea: Brachycera in Cretaceous Amber, Part VIII". American Museum Novitates. 3865 (3865): 1–29. doi:10.1206/3865.1. ISSN 0003-0082. S2CID 89602433.
  2. B. B. Rohdendorf. 1938. Dvukrylye basekomye Mezozoya Kara-Tau. I. Brachycera i chast' Nematocera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7:29-67
  3. Wang, Fengyan; Shih, Chungkun; Ren, Dong; Wang, Yongjie (January 2017). "Quantitative assessments and taxonomic revision of the genus Archirhagio with a new species from Daohugou, China (Diptera: Archisargidae)". Systematic Entomology. 42 (1): 230–239. doi:10.1111/syen.12204. S2CID 90656127.
  4. J. Zhang and H. Zhang. 2003. Two new species of archisargids (Insecta: Diptera: Archisargidae) from the Upper Jurassic Daohugou Formation (Inner Mongolia, northeastern China). Paleontological Journal 37:409-412
  5. Zhang, Junfeng (2015-10-03). "Archisargoid flies (Diptera, Brachycera, Archisargidae and Kovalevisargidae) from the Jurassic Daohugou biota of China, and the related biostratigraphical correlation and geological age". Journal of Systematic Palaeontology. 13 (10): 857–881. doi:10.1080/14772019.2014.960902. ISSN 1477-2019. S2CID 84512383.
  6. M. B. Mostovski. 1997. On knowledge of fossil flies of the superfamily Archisargoidea (Diptera, Brachycera). Paleontological Journal 31(1):72-78
  7. K. Y. Zhang, D. Yang, D. Ren and C. K. Shih. 2007. The earliest species of the extinct genus Archisargus from China (Diptera: Brachycera: Archisargidae). Annales Zoologici 57:827-832
  8. Zhang, Kuiyan; Yang, Ding; Ren, Dong; Shih, Chungkun (2007-11-23). "The oldest Calosargus Mostovski, 1997 from the Middle Jurassic of China (Diptera: Brachycera: Archisargidae)". Zootaxa. 1645 (1): 1–17. doi:10.11646/zootaxa.1645.1.1. ISSN 1175-5334.
  9. Oberprieler, Stefanie K.; Yeates, David K. (July 2012). "Calosargus talbragarensis new species: the first brachyceran fly from the Jurassic of Australia (Diptera, Archisargidae)". Journal of Paleontology. 86 (4): 641–645. doi:10.1666/11-126R.1. ISSN 0022-3360. S2CID 128737218.
  10. Zhang, Jun Feng (March 2010). "Records of bizarre Jurassic brachycerans in the Daohugou biota, China (Diptera, Brachycera, Archisargidae and Rhagionemestriidae)". Palaeontology. 53 (2): 307–317. doi:10.1111/j.1475-4983.2010.00934.x.
  11. Zhang, Junfeng (September 2012). "Distinct but rare archisargid flies from Jurassic of China (Diptera, Brachycera, Archisargidae) with discussion of the systematic position of Origoasilus pingquanensis Zhang et al., 2011". Journal of Paleontology. 86 (5): 878–885. doi:10.1666/12-028.1. ISSN 0022-3360. S2CID 86217269.
  12. Y. C. Hong. 1983. Middle Jurassic Fossil Insects in North China 1-223
  13. M. B. Mostovski. 1996. New Species of the Genus Mesosolva Hong, 1983 (Diptera, Archisargidae) from the Jurassic of Kazakhstan and Mongolia. The Continental Jurassic 60:329-332
  14. Zhang, Kuiyan; Yang, Ding; Ren, Dong; Shih, Chungkun (March 2010). "New archisargids from China (Insecta: Diptera)". Entomological Science. 13 (1): 75–80. doi:10.1111/j.1479-8298.2009.00350.x. S2CID 84716263.
  15. Junfeng, Zhang (2012-11-06). "Orientisargidae fam. n., a new Jurassic family of Archisargoidea (Diptera, Brachycera), with review of Archisargidae from China". ZooKeys (238): 57–76. doi:10.3897/zookeys.238.3624. ISSN 1313-2970. PMC 3496945. PMID 23226708.
  16. K. Y. Zhang, D. Yang, and D. Ren. 2011. A new brachyceran family Origoasilidae Fam. Nov. from the late Mesozoic of China (Insecta: Diptera). Acta Geologica Sinica 85(5):994-997
  17. M. B. Mostovski. 1996. To the knowledge of Archisargoidea (Diptera, Brachycera). Families Eremochaetidae and Archisargidae. Russian Entomological Journal 5:117-124
  18. Zhang, Kuiyan; Yang, Ding; Ren, Dong (June 2008). "Middle Jurassic fossils of the genus Sharasargus from Inner Mongolia, China (Diptera: Archisargidae)". Entomological Science. 11 (2): 269–272. doi:10.1111/j.1479-8298.2008.00271.x. S2CID 84139157.
  19. D. Ren. 1998. Late Jurassic Brachycera from northeastern China (Insecta: Diptera). Acta Zootaxonomica Sinica 23:65-82
  20. Zhang, Kuiyan; Yang, Ding; Ren, Dong; Shih, Chungkun (March 2010). "An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera): FOSSIL FLIES FROM THE MIDDLE JURASSIC OF CHINA". Zoological Journal of the Linnean Society. 158 (3): 563–572. doi:10.1111/j.1096-3642.2009.00552.x.
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