Scythris

Scythris is a genus of gelechioid moths. It is the type genus of the flower moth family, which is sometimes included as a subfamily in the Xyloryctidae, or together with these merged into the Oecophoridae.[2] The genus was erected by Jacob Hübner in 1825.

Scythris
Imago of Scythris heinemanni at Cerreto Ratti in Borghetto di Borbera, Italy
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
Division:	Ditrysia
Family:	Scythrididae (disputed)
Genus:	Scythris Hübner, [1825]
Type species
Tinea chenopodiella Hübner, [1813]
Species
Numerous, see text
Synonyms[1]
Butalis Treitschke, 1833 Copida Sodoffsky, 1837 Galanthia Hübner, 1825 Rubioia Agenjo, 1962

It is the largest genus of flower moths, and as such might not be fully monophyletic with regard to some very small or monotypic genera placed in the same family. In addition, new species of Scythris continue to be discovered and described.[3]

Selected species

Species groups

The species of Scythris have been divided among several groups, which may or may not be monophyletic; those that are may – provided they are closely enough related to the type species S. limbella – be considered subgenera. There is a considerable number of species whose exact relationships are hitherto elusive. Some have been placed in a "Monospecific species group" of their own, as they are too distinct from the other groups to be included there, yet at the same time have characteristic autapomorphies. Species groups include:[4][3]

aenea group Scythris aenea Passerin d'Entrèves 1984 aerariella group Scythris adustella Jäckh, 1978 Scythris aerariella (Herrich-Schäffer, 1855) Scythris anomaloptera (Staudinger, 1880) Scythris binotiferella (Ragonot, 1880) Scythris brummanae Passerin d'Entrèves & Roggero, 2012 Scythris carboniella Jäckh, 1978 Scythris corsa Passerin d'Entrèves, 1986 Scythris dorycniella (Milliére, 1861) Scythris flaviventrella (Herrich-Schäffer, 1855) Scythris hungaricella Rebel, 1917 Scythris imperiella Jäckh, 1978 Scythris jaeckhi Bengtsson, 1989 Scythris lhommei Bengtsson & Passerin d'Entrèves, 1988 Scythris parnassiae Bengtsson, 1997 Scythris popescugorji Passerin d'Entrèves, 1984 Scythris ridiculella Caradja, 1920 Scythris tergestinella (Zeller, 1855) Scythris tumidella K. & T.Nupponen, 2001 alseriella group Scythris alseriella (Turati, 1879) Scythris fasciatella (Ragonot, 1880) Australian group Scythris adelopa Meyrick, 1897 Scythris celidopa Meyrick, 1921 Scythris ceratocosma Meyrick, 1897 Scythris crypsigramma Meyrick, 1897 Scythris detestata Meyrick, 1922 Scythris diatoma (Turner, 1927) Scythris erebospila Meyrick, 1897 Scythris fumida Turner, 1923 Scythris hologramma (Lower, 1899) Scythris leucochyta (Turner, 1947) Scythris paredra Meyrick, 1897 Scythris pleonectis Meyrick, 1897 Scythris plocanota Meyrick, 1897 Scythris praestructa Meyrick, 1922 Scythris rhabducha Meyrick, 1897 Scythris sporadica Meyrick, 1897 Scythris xenonympha (Lower, 1900) bagdadiella group Scythris bagdadiella Amsel, 1949 Scythris caballoides Nupponen, 2009 Scythris kabulella Passerin d'Entrèves & Roggero, 2011 Scythris semifascia Nupponen, 2010 Scythris tugaiensis Nupponen, 2009 bazaensis group Scythris bazaensis Bengtsson, 1997 boseanella group Scythris boseanella Klimesch, 1986 camelella group Scythris camelella Walsingham, 1907 canescens group Scythris angustella Nupponen, 2009 Scythris appendicella Bengtsson, 1997 Scythris articulatella Chrétien, 1915 Scythris barbatella Chrétien, 1915 Scythris biskraensis Bengtsson, 1997 Scythris canescens (Staudinger, 1880) Scythris cupellella Bengtsson, 1997 Scythris curlettii Bengtsson, 1997 Scythris deserticola Nupponen, 2010 Scythris eremella Chrétien, 1915 Scythris eucharis Walsingham, 1907 Scythris falkovitshi Nupponen, 2009 Scythris karvoneni Nupponen, 2010 Scythris maculosa Bengtsson, 1997 Scythris neftae Bengtsson, 1997 Scythris pura Walsingham, 1907 Scythris senecai Bengtsson, 1997 Scythris tessulatella Rebel, 1903 caramani group Scythris albisaxella K. & T.Nupponen, 2000 Scythris arkaimensis Bengtsson, 2000 Scythris caramani Staudinger, 1880 Scythris cervella K. & T.Nupponen, 2001 Scythris cultelloides Nupponen & Sinev, 2011 Scythris eevae Nupponen, 2007 Scythris erinacella Nupponen, 2003 Scythris gorbunovi Nupponen, 2003 Scythris hamatella K. & T.Nupponen, 2001 Scythris hebesella Nupponen, 2005 Scythris kullbergi Bengtsson, 1997 Scythris lagomorphella Nupponen, 2002 Scythris malozemovi Nupponen, 2003 Scythris marashi Passerin d'Entrèves & Roggero, 2012 Scythris sinevi Nupponen, 2003 cicadella group Scythris arenbergeri Passerin d'Entrèves, 1986 Scythris bajanlegi Passerin d'Entrèves & Roggero, 2006 Scythris cicadella (Zeller, 1839) Scythris latilineella K. Nupponen, 2013 Scythris moldavicella Caradja, 1905 (= S. balcanica) Scythris potentillella (Zeller, 1847) Scythris ventosella Chrétien 1907 cistorum group Scythris cistorum (Milliére, 1876) Scythris nieukerkeni Bengtsson, 1989 Scythris rondaensis Bengtsson, 1997 crassiuscula group Scythris crassiuscula (Herrich-Schäffer, 1855) crypta group Scythris crypta Hannemann 1961 decrepidella group Scythris decrepidella Bengtsson, 1997 Scythris goundafae Bengtsson, 1997 dissitella group Scythris dissitella (Zeller, 1847) elenae group Scythris elenae K.Nupponen, 2000 Scythris juerivetei Nupponen, 2005 empetrella group Scythris empetrella Karsholt & Nielsen 1976 Scythris gozmanyi Passerin d'Entrèves 1986 fallacella group Scythris baldensis Passerin d'Entrèves 1979 Scythris fallacella (Schläger 1847) Scythris larzacensis Delmas, 2010 Scythris oelandicella Müller-Rutz 1922 Scythris sappadensis Bengtsson, 1992 Scythris tremalzoi Bengtsson, 1992

formicella group Scythris formicella Chrétien, 1915 Scythris kebiliensis Bengtsson, 1997 Scythris minima Bengtsson, 1997 fuscoaenea group Scythris derrai Bengtsson, 1991 Scythris dissimilella (Herrich-Schäffer, 1855) Scythris fuscoaenea (Haworth, 1828) Scythris parafuscoaenea Bengtsson, 1991 Scythris tabelli Junnilainen, 2002 Scythris tenuivittella (Stainton, 1867) Scythris traugotti Bengtsson, 1991 Scythris vartianae Kasy, 1962 Scythris vittella (O.Costa, 1834) fuscopterella group Scythris fuscopterella Bengtsson 1977 gobiensis group Scythris gobiensis Passerin d'Entrèves & Roggero, 2006 grandipennis group Scythris bornicensis Jäckh, 1977 Scythris constanti Walsingham, 1898 Scythris cupreella (Staudinger, 1859) Scythris ericetella (Heinemann, 1872) Scythris grandipennis (Haworth, 1828) Scythris kasyi Hannemann, 1962 Scythris maroccensis Jäckh, 1977 Scythris nevadensis Passerin d'Entrèves, 1990 Scythris salviella Meess, 1910 Scythris scipionella (Staudinger, 1859) Scythris scorpionella Jäckh, 1977 Scythris siculella Jäckh, 1977 Scythris tabidella (Herrich-Schäffer, 1855) gravatella group Scythris gravatella (Zeller, 1847) iagella group Scythris iagella Chrétien 1925 inclusella group Scythris inclusella (Lederer, 1855) inertella group Scythris ethmiella Amsel, 1974 Scythris inertella (Zeller, 1855) Scythris xanthopygella (Staudinger, 1859) inspersella group Scythris inspersella (Hübner, [1817]) insulella ("ericivorella") group Scythris ericivorella (Ragonot, 1880) Scythris insulella (Staudinger, 1859) karinae group Scythris karinae Bengtsson 1991 karsholti group Scythris karsholti Bengtsson, 1997 klimeschi group Scythris guimarensis Bengtsson, 1997 Scythris klimeschi Passerin d'Entrèves, 1983 knochella group Scythris ambustella Bengtsson, 1997 Scythris aspromontis Jäckh, 1978 Scythris ciliatella Zerny, 1936 Scythris heinemanni Scythris iberica Jäckh, 1978 Scythris knochella (Fabricius, 1794) Scythris reticulella Nupponen, 2010 Scythris schawerdae Rebel, 1931 Scythris setiella (Zeller, 1870) Scythris staudingeri Jäckh, 1978 laminella group Scythris andersi Bengtsson, 1991 Scythris bolognella Jäckh, 1978 Scythris braschiella (O.Hofmann, 1897) Scythris bubaniae Walsingham, 1907 Scythris danilevskyi Nupponen & Sinev, 2011 Scythris eberhardi Bengtsson, 1997 Scythris frankeniella Chrétien, 1916 Scythris kailai Bengtsson, 1997 Scythris laminella (Denis & Schiffermüller, 1775) Scythris saxella Bengtsson, 1991 Scythris thomisioides Bengtsson, 1997 limbella group – possibly subgenus Scythris Scythris elegantella (D.Lucas, 1955) Scythris limbella lobella group Scythris lobella K. Nupponen, 2013 mariannae group Scythris mariannae (Bengtsson, 1997) martini group Scythris bengtbengtssoni Vives, 1994 Scythris martini Bengtsson, 1991 meanderis group Scythris meanderis Bengtsson, 1997 monochreella group Scythris monochreella (Ragonot, 1896) mus group Scythris mus Walsingham, 1898 nigrella group Scythris nigrella Jäckh, 1978 nipholecta group Scythris nipholecta Meyrick, 1924 noricella group Scythris noricella (Zeller, 1843) obscurella group – possibly subgenus Galanthia Scythris amphonycella (Geyer, [1836]) Scythris bengtssoni Patocka & Liska, 1989 Scythris cuspidella (Denis & Schiffermüller, 1775) Scythris flavilaterella (Fuchs, 1886) Scythris hornigii (Zeller, 1855) Scythris lampyrella (Constant, 1865) Scythris mediella (Constant, 1855) Scythris obscurella (Scopoli, 1763) Scythris rouxella (Constant 1865) (sometimes in C. hornigii, tentatively placed here) Scythris speyeri (Heinemann & Wocke, 1876) Scythris shafferi Bengtsson, 2005 Scythris abruptella Sachkov, 2013 palustris group Scythris palustris (Zeller, 1855)

pascuella group Scythris aciella Bengtsson, 1997 Scythris albidella (Stainton, 1867) Scythris albosuffusella Nupponen, 2007 Scythris barguzinensis Bengtsson & Liska, 1996 Scythris bidzilyai Sachkov, 2013 Scythris bifissella (O.Hofmann, 1889) Scythris caroxylella Falkovitsh, 1969 Scythris claudiae Passerin d'Entrèves & Roggero, 2011 Scythris cornuella Bengtsson, 1997 Scythris cramella Nupponen, 2010 Scythris flavidella Preissecker, 1911 Scythris fluxilis Falkovitsh, 1986 Scythris hemicycliella Nupponen, 2005 Scythris kantarae Bengtsson, 1997 Scythris karinupponei Bengtsson, 2000 Scythris lagunae Jäckh, 1978 Scythris mikkolai Sinev, 1993 Scythris nigridorsella Nupponen, 2007 Scythris orientella Sinev, 2001 Scythris pascuella (Zeller, 1855) Scythris paullella (Herrich-Schäffer, 1855) Scythris perlucidella K. & T.Nupponen, 2000 Scythris pudorinella (Möschler, 1866) Scythris rotundella Nupponen, 2010 Scythris satyrella Staudinger, 1880 Scythris subaerariella (Stainton, 1867) Scythris tabescentella (Staudinger, 1880) passerini group Scythris passerini (Bengtsson, 1997) penicillata group Scythris buraetica Nupponen, 2007 Scythris penicillata (Chrétien 1900) Scythris spinella K. & T.Nupponen, 2001 petrella group Scythris arachnodes Walsingham, 1908 Scythris hierroella Klimesch, 1986 Scythris petrella Walsingham, 1908 Scythris pseudoarachnodes Bengtsson, 1997 picaepennis group Scythris disparella (Tengström, 1848) Scythris picaepennis (Haworth, 1828) pinker group Scythris pinkeri Klimesch, 1986 platypyga group Scythris platypyga (Staudinger, 1880) podoliensis group Scythris podoliensis Rebel, 1938 polycarpaeae group Scythris polycarpaeae Klimesch, 1986 productella group Scythris nielseni Passerin d'Entrèves & Roggero, 2003 Scythris productella (Zeller, 1839) pulicella group Scythris antisymmetrica Nupponen, 2009 Scythris apotomella Nupponen, 2007 Scythris azrouensis Bengtsson, 1997 Scythris corleyi Bengtsson, 1997 Scythris langohri Passerin d'Entrèves, 1990 Scythris pilella Bengtsson, 1991 Scythris pulicella (Staudinger, 1859) Scythris skulei Bengtsson, 1997 Scythris veletae Passerin d'Entrèves, 1990 punctivittella group Scythris albostriata Hannemann, 1961 Scythris apicistrigella (Staudinger, 1870) Scythris atlasensis Bengtsson, 1997 Scythris confluens (Staudinger, 1870) Scythris cycladeae Jäckh, 1978 Scythris emichi (Anker, 1870) Scythris landryi Passerin d'Entrèves & Roggero, 2003 Scythris punctivittella (O.Costa, 1836) Scythris trinacriae Passerin d'Entrèves 1984 rubioi group Scythris rubioi Agenjo, 1962 – possibly subgenus Rubioia schleichiella group Scythris glacialis (Frey, 1870) Scythris gratiosella Jäckh, 1978 Scythris schleichiella (Zeller, 1870) Scythris similis Hannemann, 1961 Scythris subschleichiella Hannemann, 1961 scopolella group Scythris apicalis (Zeller, 1847) Scythris flabella (Mann, 1861) Scythris scopolella seliniella group Scythris acarioides Bengtsson, 1997 Scythris clavella (Zeller, 1855) Scythris seliniella (Zeller, 1839) Scythris subseliniella (Heinemann, 1876) Scythris villari Agenjo, 1971 siccella group Scythris lafauryi Passerin d'Entrèves 1986 Scythris lempkei Bengtsson & Langohr, 1989 Scythris siccella (Zeller, 1839) Scythris subsiccella Bengtsson, 1997 sinensis group Scythris sinensis (Felder & Rogenhofer, 1875) subfasciata group Scythris anthracodes Walsingham, 1907 Scythris friedeli Bengtsson, 1997 Scythris levantina Passerin d'Entrèves & Vives, 1990 Scythris locustella Chrétien, 1915 Scythris pototskii Nupponen, 2005 Scythris pseudolocustella Passerin d'Entrèves & Vives, 1990 Scythris subfasciata (Staudinger, 1880) tributella group Scythris tributella (Zeller, 1847) unquisella group Scythris unquisella Bengtsson, 2005 ? species group Scythris herati Passerin d'Entrèves & Roggero, 2003 Scythris tauromeniella Passerin d'Entrèves & Roggero, 2003

Species incertae sedis

Finally, there are many species which are neither clearly assignable to any one species group, nor autapomorphic enough to be included in a monospecific "group". These include:[3]

Scythris abyanensis Bengtsson, 2002 Scythris accumulata Meyrick, 1914 Scythris achyropa Meyrick, 1916 Scythris acipenserella K. & T.Nupponen, 2000 Scythris acusella Passerin d'Entrèves & Roggero, 2012 Scythris adelopa Meyrick, 1897 Scythris aegrella K. Nupponen & Junnilainen, 2000 Scythris albiangulella Bengtsson, 2002 Scythris albipuncta Turner, 1939 Scythris albocanella Bengtsson, 2002 Scythris albogrammella Bengtsson, 2002 Scythris alborzensis Passerin d'Entrèves & Roggero, 2011 Scythris alceella Junnilainen, 2002 Scythris alhamrae Bengtsson, 2002 Scythris amplexella Bengtsson, 2002 Scythris ampullella Passerin d'Entrèves & Roggero, 2012 Scythris anchophylli Falkovitsh, 1969 Scythris ancystra Bucheli, 2005 Scythris anthracina Braun, 1923 Scythris anthracodelta Meyrick, 1911 Scythris aquaria Meyrick, 1913 Scythris arenicola K.Nupponen, 2005 Scythris arerai Huemer, 2000 Scythris aristidella Rebel, 1902 Scythris astragali Falkovitsh, 1969 Scythris atascosa (Landry, 1991) Scythris atollicola Nupponen & Saldaitis, 2013 Scythris autochlorella Paulian & Viette, 1955 Scythris axenopa Meyrick, 1918 Scythris balanophora Meyrick, 1916 Scythris balkhi Passerin d'Entrèves & Roggero, 2008 Scythris balli (Landry, 1991) Scythris basilaris (Zeller, 1855) Scythris basilicella Bengtsson, 2002 Scythris basistrigella (Staudinger, 1880) Scythris beccella Bengtsson, 2002 Scythris bernardlandryi Bucheli, 2005 Scythris biacutella Bengtsson, 2002 Scythris bicuspidella Bengtsson, 2002 Scythris bifasciella Passerin d'Entrèves & Roggero, 2011 Scythris bifractella (Rebel, 1917) Scythris bispinella Bengtsson, 2002 Scythris brachyplecta Meyrick, 1928 Scythris brandti Passerin d'Entrèves & Roggero, 2008 Scythris brunneofasciella K. Nupponen & Junnilainen, 2000 Scythris buszkoi Baran, 2003 Scythris canceroides Bengtsson, 1997 Scythris canella Bengtsson, 2002 Scythris capitalis (Erschoff, 1874) Scythris capnofasciae Bengtsson, 2002 Scythris cassiterella (Snellen, 1884) (= S. baikalensis) Scythris celidopa Meyrick, 1921 Scythris ceratella Bengtsson, 2002 Scythris ceratocosma Meyrick, 1897 Scythris charon Meyrick, 1918 Scythris chelota Meyrick, 1906 Scythris chloraema (Meyrick, 1887) Scythris chrysopygella Caradja, 1927 Scythris cinisella Bengtsson, 2002 Scythris cirra Falkovitsh, 1969< Scythris clemens Meyrick, 1921 Scythris cometa Meyrick, 1909 Scythris commota Meyrick, 1918 Scythris complexa Sinev, 2001 Scythris concurrens Meyrick, 1921 Scythris consimilella Bengtsson, 2002 Scythris cretacella K. & T.Nupponen, 2000 Scythris cretiflua Meyrick, 1913 Scythris crypsigramma Meyrick, 1897 Scythris cucullella Bengtsson, 2002 Scythris culmicola Meyrick, 1916 Scythris cuneata Bucheli, 2005 Scythris cuneatella Bengtsson, 2002 Scythris curvipilella Bengtsson, 2002 Scythris darwini Bucheli, 2005 Scythris delodelta Meyrick, 1930 Scythris denticolor Walsingham, 1900 Scythris deplanata Meyrick, 1928 Scythris depressa Meyrick, 1931 Scythris deprinsi Bengtsson, 2005 Scythris deresella Falkovitsh, 1969 Scythris detestata Meyrick, 1922 Scythris diatoma (Turner, 1927) Scythris dicroa Falkovitsh, 1972 Scythris digitibasella Nupponen & Saldaitis, 2013 Scythris dimensa Meyrick, 1920 Scythris dimota Meyrick, 1931 Scythris discimaculella Rebel, 1936 Scythris distactica Meyrick, 1921 Scythris divaricata Braun, 1923 Scythris divergens Bengtsson, 2005 Scythris dividua Meyrick, 1916 Scythris durandella (Lucas, 1949) Scythris eboracensis (Zeller, 1855) Scythris eburnea (Walsingham, 1888) Scythris ejiciens Meyrick, 1928

Scythris elachistoides Bengtsson, 2002 Scythris elburzi Passerin d'Entrèves & Roggero, 2008 Scythris eloquens Meyrick, 1921 Scythris epistrota (Meyrick, 1889) Scythris erebospila Meyrick, 1897 Scythris erinacella Nupponen, 2003 Scythris eversmanni K. & T. Nupponen, 2000 Scythris expolita Meyrick, 1910 Scythris exsoluta Meyrick, 1920 Scythris faeculenta Meyrick, 1912 Scythris falcata Bucheli, 2005 Scythris farrata Meyrick, 1913 Scythris farsi Passerin d'Entrèves & Roggero, 2009 Scythris fasciata Falkovitsh, 1969 Scythris felesella Bengtsson, 1997 Scythris felixi Bengtsson & Sutter, 1996 Scythris fibigeri Bengtsson, 2002 Scythris fissurella Bengtsson, 1996 Scythris fluvialis Meyrick, 1916 Scythris fonticola Meyrick, 1911 Scythris formidabilis (Landry, 1991) Scythris fumida Turner, 1923 Scythris furculata Bucheli, 2005 Scythris fuscicomella (Clemens, 1860) Scythris galapagosensis Bucheli, 2005 Scythris galeatella Bengtsson, 2002 Scythris ganesha Nupponen & Sinev, 2011 Scythris garciapitai Vives, 2001 Scythris ghaemii Bengtsson & Huemer, 2003 Scythris gladiella K. & T.Nupponen, 2004 Scythris glaphyropa Meyrick, 1914 Scythris glyphidota Meyrick, 1916 Scythris gratifica Meyrick, 1921 Scythris halmyrodes Meyrick, 1921 Scythris halophiliella Amsel, 1935 Scythris haloxylella Falkovitsh, 1969 Scythris hemidictyas Meyrick, 1928 Scythris hamardabanica Nupponen, 2003 Scythris heikkii Nupponen, 2007 Scythris hologramma (Lower, 1899) Scythris hostilis Nupponen, 2005 Scythris hyalinella Caradja, 1920 Scythris hydronoma Meyrick, 1930 Scythris hymenata (Landry, 1991) Scythris ianitella Nupponen, 2005 Scythris iconiensis Rebel, 1902 Scythris ilyopa Meyrick, 1921 Scythris immaculatella (Chambers, 1875) Scythris immunis Meyrick, 1916 Scythris inanima Meyrick, 1916 Scythris inconspicuella Sinev, 2001 Scythris indigoferivora Bengtsson, 2002 Scythris ingens Nupponen, 2005 Scythris inornatella (Chambers, 1880) Scythris interrupta Braun, 1920 Scythris invisa Meyrick, 1920 Scythris iterella Bengtsson, 2002 Scythris jakutica Sinev, 2001 Scythris jalavai Sinev, 1993 Scythris jemenensis Bengtsson, 2002 Scythris justifica Meyrick, 1911 Scythris karini Bengtsson, 1991 Scythris kaszabi Passerin d'Entrèves & Roggero, 2006 Scythris kaschmirica Bengtsson, 2005 Scythris kefensis Nupponen, 2001 Scythris kolachii Passerin d'Entrèves & Roggero, 2012 Scythris kyzylensis Bengtsson, 1997 Scythris lactanea Meyrick, 1913 Scythris lamprochalca Meyrick, 1931 Scythris latebrosa Meyrick, 1913 Scythris lativalvella Sinev, 2001 Scythris lativittis Gerasimov, 1930 Scythris leucochyta (Turner, 1947) Scythris libanotica Jäckh, 1978 Scythris linnavuorii Bengtsson, 2005 Scythris longissima (Landry, 1991) Scythris luxatiella K.Nupponen & Kaitila, 2000 Scythris lycii Falkovitsh, 1969 Scythris macrourella Sinev, 2001 Scythris medullata Meyrick, 1916 Scythris melanodora Meyrick, 1912 Scythris meraula Meyrick, 1916 Scythris mesoplecta Meyrick, 1921 Scythris minorella Sinev, 2001 Scythris mixaula Meyrick, 1916 Scythris nanophyti Falkovitsh, 1979 Scythris neocompsa (Meyrick, 1933) Scythris nepalensis Bengtsson, 2005 Scythris neurogramma Walsingham, 1900 Scythris nigra Philpott, 1931 Scythris nigrispersa Meyrick, 1918 Scythris nigrogrammella Bengtsson, 2002 Scythris nigropterella Bengtsson, 2002 Scythris ninae Nupponen, 2003 Scythris niphozela Meyrick, 1931 Scythris nitidella Bengtsson & Liska, 1996 Scythris nivicolor Meyrick, 1916 Scythris notorrhoa Meyrick, 1921 Scythris obliqua Falkovitsh, 1969 Scythris ochrantha Meyrick, 1909 Scythris ochrea Walsingham, 1896 Scythris ochrogramma Meyrick, 1916 Scythris olschwangi K. & T.Nupponen, 2000 Scythris onerica Nupponen, 2009

Scythris orientella Sinev, 2001 Scythris oxyplecta Meyrick, 1916 Scythris paelopyga Staudinger, 1880 Scythris pallidella Passerin d'Entrèves & Roggero, 2006 Scythris pamirica Passerin d'Entrèves & Roggero, 2008 Scythris pangalactis Meyrick, 1933 Scythris parachalca Meyrick, 1916 Scythris parafluxilis Passerin d'Entrèves & Roggero, 2007 Scythris paralogella Bengtsson, 2002 Scythris paredra Meyrick, 1897 Scythris parenthesella Bengtsson, 2002 Scythris patiens Meyrick, 1921 Scythris pectinicornis Walsingham, 1900 Scythris pelinaula Meyrick, 1916 Scythris pelochyta Meyrick, 1909 Scythris persephone Meyrick, 1936 Scythris pfeifferella Rebel, 1936 Scythris pilosella (Zeller, 1873) Scythris pistillata Bucheli, 2005 Scythris piratica Meyrick, 1928 Scythris pleonectis Meyrick, 1897 Scythris plocanota Meyrick, 1897 Scythris plocogastra Meyrick, 1931 Scythris plutelloidella Turati, 1934 Scythris poliantha Meyrick, 1921 Scythris pollicella Bengtsson, 2002 Scythris potatorella Nupponen, 2003 Scythris powelli (Landry, 1991) Scythris praematura Meyrick, 1937 Scythris praestructa Meyrick, 1922 Scythris pruinata Falkovitsh, 1972 Scythris pterosaurella Bengtsson, 2002 Scythris reflectella Bengtsson, 2002 Scythris remexella K. Nupponen & Kaitila, 2000 Scythris rhabducha Meyrick, 1897 Scythris richteri Bengtsson, 2013 Scythris rivigera Meyrick, 1911 Scythris roseola Meyrick, 1912 Scythris saarelai K. & T.Nupponen, 1999 Scythris sacharissa Meyrick, 1913 Scythris sachkovi Passerin d'Entrèves & Roggero, 2006 Scythris sagitatella Erschoff, [1877] Scythris sanae Bengtsson, 2002 Scythris sciochalca Meyrick, 1928 Scythris scotinopa Meyrick, 1935 Scythris scutulella Nupponen, 2013 Scythris scyphella Bengtsson, 2002 Scythris sibirella Sinev, 2001 Scythris sinuosa Bucheli, 2005 Scythris sinuosella Bengtsson, 2002 Scythris sitarcha Meyrick, 1918 Scythris strabella Bengtsson, 2002 Scythris solitaria Diakonoff, 1955 Scythris soluta Meyrick, 1916 Scythris sophronia Meyrick, 1935 Scythris sordidella Bengtsson, 2002 Scythris spectatorella Nupponen, 2001 Scythris spissata Meyrick, 1916 Scythris sponsella (Busck, 1907) Scythris sporadica Meyrick, 1897 Scythris spumifera Meyrick, 1918 Scythris stagnosa Meyrick, 1913 Scythris striella Turati, 1929 Scythris subcassiterella Bengtsson, 1997 Scythris subclavella Rebel, 1901 Scythris subeburnea (Walsingham, 1891) Scythris subgaleatella Bengtsson, 2002 Scythris sublaminella K. & T.Nupponen, 2000 Scythris subparachalca Bengtsson, 2002 Scythris subsignata Meyrick, 1916 Scythris syrmatica Meyrick, 1916 Scythris taizzae Bengtsson, 2002 Scythris talyniella Passerin d'Entrèves & Roggero, 2006 Scythris taurella Caradja, 1920 Scythris tauromeniella Passerin d'Entrèves & Roggero, 2004 Scythris taygeticola Scholz, 1997 Scythris tenuisquamata Staudinger, 1880 Scythris tephrella Bengtsson, 2005 Scythris terekholensis Bengtsson, 1997 Scythris tibicina Meyrick, 1916 Scythris timoi Nupponen, 2009 Scythris triatma Meyrick, 1936 Scythris trinummulata Meyrick, 1922 Scythris triochrias Meyrick, 1916 Scythris trivinctella (Zeller, 1873) Scythris tsherkesella Falkovitsh, 1969 Scythris turanica Nupponen, 2009 Scythris tuzensis Bengtsson, 2005 Scythris tytrella Falkovitsh, 1969 Scythris unimaculella Rebel, 1905 Scythris ustjuzhanini Sachkov & Sinev, 2001 Scythris valgella Bengtsson, 2002 Scythris valvaearcella Bengtsson, 2002 Scythris vernusella Jäckh, 1978 Scythris vogelfederbergensis (Mey, 2011) Scythris xenonympha (Lower, 1900) Scythris xylinochra Meyrick, 1931 Scythris ypsilon Braun, 1920 Scythris wadiqeltella Passerin d'Entrèves & Roggero, 2013 Scythris zelleri (Staudinger, 1880) Scythris zeugmatica Meyrick, 1931

Afrotropical

Scythris aarviki Bengtsson, 2014 Scythris abachausensis Bengtsson, 2014 Scythris agassizi Bengtsson, 2014 Scythris alainensis Bengtsson, 2014 Scythris albipunctella Bengtsson, 2014 Scythris albonigrella Bengtsson, 2014 Scythris anaecapitella Bengtsson, 2014 Scythris apicispinella Bengtsson, 2014 Scythris aratrella Bengtsson, 2014 Scythris asinella Bengtsson, 2014 Scythris atroparvella Bengtsson, 2014 Scythris aulaeella Bengtsson, 2014 Scythris balantiella Bengtsson, 2014 Scythris baringensis Bengtsson, 2014 Scythris basimaculella Bengtsson, 2014 Scythris bernardi Bengtsson, 2014 Scythris bicalamella Bengtsson, 2014 Scythris bipunctella Bengtsson, 2014 Scythris bisacculella Bengtsson, 2014 Scythris bisincusella Bengtsson, 2014 Scythris bitterfonteinica Bengtsson, 2014 Scythris bjoernstadi Bengtsson, 2014 Scythris bontebokensis Bengtsson, 2014 Scythris bosicornella Bengtsson, 2014 Scythris brandbergensis Bengtsson, 2014 Scythris brevimanubriella Bengtsson, 2014 Scythris bromiella Bengtsson, 2014 Scythris brunneostriella Bengtsson, 2014 Scythris budongensis Bengtsson, 2014 Scythris capilliverticella Bengtsson, 2014 Scythris catuliformis Bengtsson, 2014 Scythris cederbergensis Bengtsson, 2014 Scythris clarki Bengtsson, 2014 Scythris claudioculella Bengtsson, 2014 Scythris conimarginella Bengtsson, 2014 Scythris coniobliquella Bengtsson, 2014 Scythris cooperi Bengtsson, 2014 Scythris coriella Bengtsson, 2014 Scythris cottrelli Bengtsson, 2014 Scythris cricetinaeformis Bengtsson, 2014 Scythris davidi Bengtsson, 2014 Scythris dicksoni Bengtsson, 2014 Scythris dorsifuscella Bengtsson, 2014 Scythris durbanensis Bengtsson, 2014 Scythris eburiplicella Bengtsson, 2014 Scythris eburnella Bengtsson, 2014 Scythris eburnipterella Bengtsson, 2014 Scythris ellipsiella Bengtsson, 2014 Scythris enigmella Bengtsson, 2014 Scythris ethiopica Bengtsson, 2014 Scythris etoshensis Bengtsson, 2014 Scythris falciformis Bengtsson, 2014 Scythris flavoterminella Bengtsson, 2014

Scythris fumarolella Bengtsson, 2014 Scythris gaboronensis Bengtsson, 2014 Scythris geminella Bengtsson, 2014 Scythris gielisi Bengtsson, 2014 Scythris gilgilensis Bengtsson, 2014 Scythris griseella Bengtsson, 2014 Scythris hanseni Bengtsson, 2014 Scythris helskloofensis Bengtsson, 2014 Scythris heniaeguttella Bengtsson, 2014 Scythris hermanusensis Bengtsson, 2014 Scythris hirudoformis Bengtsson, 2014 Scythris humeriformis Bengtsson, 2014 Scythris jacobseni Bengtsson, 2014 Scythris jamakensis Bengtsson, 2014 Scythris jansei Bengtsson, 2014 Scythris jodhpursoides Bengtsson, 2016 Scythris jurateae Bengtsson, 2014 Scythris kalaharii Bengtsson, 2014 Scythris kalkrandensis Bengtsson, 2014 Scythris kavangensis Bengtsson, 2014 Scythris kihondensis Bengtsson, 2014 Scythris kilifiensis Bengtsson, 2014 Scythris kingstoni Bengtsson, 2014 Scythris krooni Bengtsson, 2014 Scythris kruegeri Bengtsson, 2014 Scythris lactifuscella Bengtsson, 2014 Scythris lahaivora Bengtsson, 2014 Scythris leifi Bengtsson, 2014 Scythris lushotensis Bengtsson, 2014 Scythris magnipedella Bengtsson, 2014 Scythris malawica Bengtsson, 2014 Scythris malelanensis Bengtsson, 2014 Scythris marginifuscella Bengtsson, 2014 Scythris matopensis Bengtsson, 2014 Scythris messinensis Bengtsson, 2014 Scythris meyeri Bengtsson, 2014 Scythris meyi Bengtsson, 2014 Scythris mpalensis Bengtsson, 2014 Scythris mulanjensis Bengtsson, 2014 Scythris munroi Bengtsson, 2014 Scythris naivashensis Bengtsson, 2014 Scythris najaoides Bengtsson, 2014 Scythris naukluftensis Bengtsson, 2014 Scythris nguliae Bengtsson, 2014 Scythris niniae Bengtsson, 2014 Scythris nussi Bengtsson, 2014 Scythris nyangensis Bengtsson, 2014 Scythris nyikensis Bengtsson, 2014 Scythris nylstroomensis Bengtsson, 2014 Scythris nylsvleyensis Bengtsson, 2014 Scythris obnubilella Bengtsson, 2014 Scythris ochrocrusella Bengtsson, 2014 Scythris ochroplicella Bengtsson, 2014 Scythris octocornella Bengtsson, 2014

Scythris oculella Bengtsson, 2014 Scythris otaviensis Bengtsson, 2014 Scythris paarlensis Bengtsson, 2014 Scythris palmwagensis Bengtsson, 2014 Scythris piriensis Bengtsson, 2014 Scythris piriformis Bengtsson, 2014 Scythris popensis Bengtsson, 2014 Scythris potgieteri Bengtsson, 2014 Scythris pravitella Bengtsson, 2014 Scythris pretoriensis Bengtsson, 2014 Scythris pulveratella Bengtsson, 2014 Scythris quadrilobella Bengtsson, 2014 Scythris rumurutiensis Bengtsson, 2014 Scythris satarensis Bengtsson, 2014 Scythris scholzi Bengtsson, 2014 Scythris schouteni Bengtsson, 2014 Scythris sericiella Bengtsson, 2014 Scythris serinusoides Bengtsson, 2014 Scythris setaelongella Bengtsson, 2014 Scythris shingwedziensis Bengtsson, 2014 Scythris silfverbergi Bengtsson, 2014 Scythris simplicella Bengtsson, 2014 Scythris skukuzensis Bengtsson, 2014 Scythris snymani Bengtsson, 2014 Scythris solutella Bengtsson, 2014 Scythris somangensis Bengtsson, 2014 Scythris stoltzei Bengtsson, 2014 Scythris strydomi Bengtsson, 2014 Scythris subconcurrens Bengtsson, 2014 Scythris subcurvipilella Bengtsson, 2014 Scythris subgriseella Bengtsson, 2014 Scythris subnigropterella Bengtsson, 2014 Scythris subroseola Bengtsson, 2014 Scythris thikensis Bengtsson, 2014 Scythris thoracifaciella Bengtsson, 2014 Scythris tubulella Bengtsson, 2014 Scythris turiensis Bengtsson, 2014 Scythris ugabensis Bengtsson, 2014 Scythris ugandica Bengtsson, 2014 Scythris umtaliensis Bengtsson, 2014 Scythris unciclavella Bengtsson, 2014 Scythris valvaerimella Bengtsson, 2014 Scythris vanderwolfi Bengtsson, 2014 Scythris varii Bengtsson, 2014 Scythris virgaeformis Bengtsson, 2014 Scythris wankiensis Bengtsson, 2014 Scythris waterbergensis Bengtsson, 2014 Scythris wolframi Bengtsson, 2014 Scythris worcesterensis Bengtsson, 2014 Scythris zimbabwensis Bengtsson, 2014

Selected former species

Scythris ammobia Falkovitsh, 1972 Scythris annae Bengtsson, 1997 Scythris aphanatma Meyrick, 1933 Scythris asema Falkovitsh, 1972 Scythris asthena Falkovitsh, 1972 Scythris badiella Bengtsson, 2002 Scythris calciflua Meyrick, 1921

Scythris canispersa Meyrick, 1913 Scythris erudita Meyrick, 1917 Scythris fluctuosa Meyrick, 1914 Scythris homoxantha Meyrick, 1921 Scythris hypolepta Falkovitsh, 1972 Scythris melanopleura Meyrick, 1914

Scythris meligastra Meyrick, 1920 Scythris mongholica Passerin d'Entrèves & Roggero, 2006 Scythris pediculella Bengtsson, 1997 Scythris physalis Falkovitsh, 1972 Scythris psamathota Meyrick, 1913 Scythris zylinochra Meyrick, 1931

Footnotes

1. AEBR (2008), and see references in Savela (2008)
2. Pitkin & Jenkins (2004), AEBR (2008), ToL (2008), FE (2009), and see references in Savela (2008)
3. FE (2009), and see references in Savela (2008)
4. AEBR (2008), FE (2009), and see references in Savela (2008)

References

• Australian Biological Resources Study (ABRS) (2008): Australian Faunal Directory – Scythris. Version of 9 October 2008. Retrieved 1 May 2010.
• Fauna Europaea (FE) (2009): Scythris. Version 2.1, 22 December 2009. Retrieved 1 May 2010.
• Pitkin, Brian & Jenkins, Paul (2004): Butterflies and Moths of the World, Generic Names and their Type-species – Scythris. Version of 5 November 2004. Retrieved 1 May 2010.
• Savela, Markku (2008): Markku Savela's Lepidoptera and some other life forms – Scythris. Version of 17 July 2008. Retrieved 1 May 2010.
• Tree of Life Web Project (ToL) (2009): Xyloryctidae. Version of 1 May 2008. Retrieved 30 April 2010.