Hasle Formation

Hasle Formation
Stratigraphic range: Pliensbachian
~185 Ma

PreꞒ

Ꞓ

O

S

D

C

P

T

J

K

Pg

N

↓
Hasle Formation; Stratigraphic range: Pliensbachian; ~185 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Underlies	Sorthat Formation
Overlies	Rønne Formation
Thickness	80–140 m (260–460 ft)
Lithology
Primary	Siltstone, sandstone
Location
Coordinates	55.2°N 14.7°E
Approximate paleocoordinates	45.0°N 18.5°E
Region	Bornholm
Country	Denmark
	Hasle Formation (Denmark)

The Hasle Formation is a geologic formation on the island on Bornholm, Denmark. It is of early to late Pliensbachian age. Vertebrate fossils have been uncovered from this formation.[1][2]

Fossil content

Unidentified fin spines are known from this formation.[1]

Genus	Species	Material	Notes
Hybodus[1]	Hybodus reticulatus[1]	21 incomplete crowns	A marine shark, type member of the family Hybodontidae inside Hybodontiformes.
Hybodus[1]	Hybodus delabechei[1]	Three complete crowns	A marine shark, type member of the family Hybodontidae inside Hybodontiformes. It´s teeth of suggest that it may have been feeding on shelly invertebrates.
Hybodontidae[1]	Hybodontidae Indeterminate	One incomplete spine	A marine shark, Incertade sedis inside Hybodontiformes.
Lissodus	Lissodus hasleensis[1]	Four complete teeth	A marine shark, member of the family Lonchidiidae inside Hybodontiformes.
Synechodus[1]	Synechodus occultidens[1]	Four incomplete teeth	A marine shark, type member of the family Paleospinacidae inside Synechodontiformes.
Paraorthacodus[1]	Paraorthacodus sp[1]	Two incomplete teeth	A marine shark, member of the family Paleospinacidae inside Synechodontiformes.
Agaleus[1]	Agaleus dorsetensis[1]	Four teeth	A marine shark, type member of the family Agaleidae inside Euselachii. This genus may have been a suction feeder as its anterior teeth are similar to those of extant nurse sharks.
Oblidens[3]	Oblidens bornholmensis[3]	Upper posterior (palatine) and lower posterior (mandibular) tooth plates.	A marine Chimaera, member of the family Myriacanthidae inside Chimaeriformes.
Myriacanthus[4][3]	Myriacanthus sp.[3]	Isolated tooth plate	A marine Chimaera, type member of the family Myriacanthidae inside Chimaeriformes.

Genus	Species	Material	Notes
Plesiosauroidea[5][6]	Plesiosauroidea Indeterminate	Tooth	A marine plesiosaur, member of the family Plesiosauroidea inside Sauropterygia.
Plesiosauria[5][6]	Plesiosauria Indeterminate	Complete right ischium Complete isolated humerus Complete ulna/fibula Isolated flattened lunate epipodial bone Fragment of jaw Sacral Vertebra Sacral Rib Single Fragment of Rib	A marine plesiosaur, member of the family Plesiosauroidea inside Sauropterygia.
Rhomaleosauridae[5][6]	Rhomaleosauridae Indeterminate	Tooth	A marine plesiosaur, member of the family Rhomaleosauridae inside Sauropterygia. Resembles Rhomaleosaurus.
Pliosauroidea[5][6]	Pliosauroidea Indeterminate	Fourteen isolated teeth	A marine plesiosaur, member of the family Pliosauroidea inside Sauropterygia. Resembles Attenborosaurus.

Genus	Species	Material	Notes	Images
Stenonyx[7][8]	Stenonyx isp.	25 mm footprints referred to a non-avian Theropod.[7]	Theropod Tracks, member of the ichnofamily Anchisauripodidae, incertade sedis inside Theropoda. Considered a very Small Dinosaur, probably a juvenile. The Foot resembles those of young Coelophysis from Ghost Ranch Quarries.[7] Stenonyx footprints had been described from the Early Jurassic Soltykow (Hettangian) and Szydlowek, what matches with specimens found on the Hasle Formation, linking Southern Sweden, Bornholm and Poland, contiguous during the Early Jurassic and dinosaurs could thus freely roam this large area.[7]

Genus	Species	Material	Notes	Images
Massopoda[9][8]	Massopoda Indeterminate	DK966, an upper part of a femur from a juvenile animal	A Massopoda Sauropodomorph. DK966 has been compared with various taxa from the lower jurassic: "Aristosaurus erectus", a young Massospondylus, various Massospondylus specimens, Lufengosaurus huenei holotype, Seitaad ruessi, Thecodontosaurus, Pantydraco, etc. DK966 can be probably a young specimen of Massopoda, related to Massospondylus, more than a Plateosaurid or an Anchisaurid. We find it related to "Hortalotarsus", "Aristosaurus" and Massospondylus more than Seitaad for example. Based on the several taxa we had compared probably presented a body similar to Pantydraco being a young specimen. The complete bone is estimated on only 6 cm, making the animal no more than 70 cm length.
Massopoda[9][8]

Genus	Species	Material	Notes
Aegoceras[2]	A. centaurus[2]	specimens[2]	Later renamed Beaniceras centaurus, senior synonym of Aegoceras centaurus[2]
	A. armatum[2]	specimens[2]
	A. capraius[2]	specimens[2]
	A. mangenesti[2]	specimens[2]
	A. pettos[2]	specimens[2]	Junior synonym of Coeloceras pettos, possible synonym of Coeloceras grenouillouxi[2]
Uncertain	? Acanthopleuroceras sp. indet.[2]	specimens[2]
Apderoceras[2]	A. aculeatum[2]	specimens[2]
Arnioceras[2]	A. falcaries[2]	specimens[2]
Beaniceras[2]	B. centaurus[2]	specimens[2]	Senior synonym of Aegoceras centaurus[2]
Coeloceras[2]	C. pettos[2]	specimens[2]	Senior synonym of Aegoceras pettos. Possible junior synonym of Coeloceras grenouillouxi[2]
Coeloceras[2]	C. grenouillouxi[2]	specimens[2]	Possible senior synonym of Coeloceras pettos[2]
Paramicroderoceras[2]	P. fila[2]	specimens[2]
Paramicroderoceras[2]	P.? sp indet[2]	specimens[2]
Phylloceras[2]	P. losconnbii[2]	specimens[2]
Phricedoceras[2]	P. taylori[2]	specimens[2]
Platypleuroceras[2]	P. brevispina[2]	specimens[2]
	P. submulticum[2]	specimens[2]
	P. caprarium[2]	specimens[2]
	P. spp. indet[2]	specimens[2]
Radstockiceras[2]	R. hechingense[2]	specimens[2]
Tragophyloceras[2]	T. numismale[2]	specimens[2]
Tragophyloceras[2]	T. sp[2]	specimens[2]
Uptonia[2]	U. lata[2]	specimens[2]
Uptonia[2]	U. sp indet[2]	specimens[2]

Rees, J. (1998). "Early Jurassic selachians from the Hasle Formation on Bornholm, Denmark" (PDF). Acta Palaeontologica Polonica. 43 (3): 43–439.
Donovan, Desmond T. & Surlyk, Finn. "Lower Jurassic (Pliensbachian) ammonites from Bornholm, Baltic Sea, Denmark" (PDF). Geological Survey of Denmark and Greenland Bulletin. 1: 555–583.CS1 maint: uses authors parameter (link)
Christopher J. Duffin; Jesper Milàn (2017). "A new myriacanthid holocephalian from the Early Jurassic of Denmark" (PDF). Bulletin of the Geological Society of Denmark. 65: 161–170.
Bonde, N., Andersen, S., Hald, N., & Jakobsen, S.L. 2008: Danekræ – Danmarks bedste fossiler, 225 pp. Gyldendal, Copenhagen.
Rees, J., & Bonde, N. (1999). Plesiosaur remains from the Early Jurassic Hasle Formation, Bornholm, Denmark. Secondary Adaptation to Life in Water, Copenhagen, 13-17.
Smith, A. S. (2008). Plesiosaurs from the Pliensbachian (Lower Jurassic) of Bornholm, Denmark. Journal of Vertebrate Paleontology, 28(4), 1213-1217.
Jesper Milàn, Finn Surlyk. 2014: An enigmatic, diminutive theropod footprint in the shallow marine Pliensbachian Hasle Formation, Bornholm, Denmark
Geocenter DK (2020):Hasle Formationen. Aviable at:https://www.geocenter.dk/geoviden/dedanskedinosaurer/hasle-formationen/
Milàn, J., Cuny, G. (2019):Knogle fra mini sauropod fundet ved Hasle. Geomuseum Faxe Danmark. University of Lyon Frankrig. https://www.researchgate.net/publication/332291953_Knogle_fra_mini_sauropod_fundet_ved_Hasle

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