Fabrosauridae

The Fabrosauridae is an obsolete group of basal ornithischian dinosaurs from the Early to Middle Jurassic, originally proposed by Galton (1972). Recent studies indicate the Fabrosaurs are not a natural grouping of dinosaurs, and instead consist of unrelated genera.[1]

Fabrosaurids Temporal range: Early–Middle Jurassic, 199–155 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Superorder:	Dinosauria
Order:	Ornithischia
Family:	Fabrosauridae
Genera
?Agilisaurus ?Fabrosaurus ?Gongbusaurus ?Lesothosaurus

Agilisaurus louderbacki

The proposed "Fabrosaurs" descended from a Lesothosaurus-like animal. Proposed fabrosaurids were 1–2 meters long, and were lightly built and bipedal. Their skulls were triangular and had very large eye sockets. They were herbivorous and would have used agility to escape predators.

Fabrosaurids are known from southern Africa in the Upper Triassic and Lower Jurassic, as well as in China from the Middle and Upper Jurassic periods.[2] Fabrosaurids bear a semblance to many other members of Ornithischia, and have attracted much interest in phylogenetic studies. However, most genera that were previously referred to as Fabrosauridae are only known from fragmented and partial remains, with most based on no more than isolated teeth, and their place in Fabrosauridae is questionable.[3]

Galton in 1972 originally proposed the creation of Fabrosauridae based on the finding of cheek teeth. The successive discovery (1990) of a Middle Jurassic primitive ornithischian, Agilisaurus louderbacki, found in the Xiashaximiao Formation of Zigong, Sichuan Basin, China, has led to paleontologists learning more about the phylogeny of fabrosaurids.

Fabrosaurids have many characters that are unique to their physiology, particularly unique tooth morphology and structure, and short forelimb lengths compared to hindlimbs.[4] Unlike many other primitive Ornithischians, such as heterodontosaurids and hypsilophodontids, the teeth of fabrosaurids are very thinly and uniformly enameled. Compared to Agilisaurus, Fabrosaurus is much more primitive in that it has six premaxillary teeth, and a stout prepubis. Agilisaurus differs from Fabrosaurus in that it appears to be larger and more well developed in structure.[5]

The Upper Jurassic ornithischian dinosaur, Gongubusaurus, from China, is thought to be more closely related to fabrosaurids than hypsilophodontids. The type species, G. shiyii, is from the Shangshaximiao Formation in the Rongxian County in Sichuan Basin, has been classified from only a few scattered teeth.[2]

Description

Fabrosaurids were relatively small ornithischian dinosaurs, averaging about 1–2 meters in body length, with a triangular skull that had large circular orbits on the sides. This suggests that fabrosaurids had relatively huge eyes that faced laterally. The forelimbs of fabrosaurids are considerably shorter than their hindlimbs. A small forelimb such as those present in Fabrosauridae would not have been useful for locomotion, and it is evident that fabrosaurids were bipedal dinosaurs. The entire skeleton was lightly built, with a largely fenestrated skull and a very stout neck and trunk. The tail is nearly half of the dinosaurs' overall length. The long tail presumably acted as a counterbalance and as a compensating mechanism for shifts in the creature's center of gravity. The hindlimbs of fabrosaurids show that the tibia is considerably longer than the femur, a feature that suggests that fabrosaurids were adapted for bipedality, and were fast runners.[6]

References

1. Weishampel, David B.; Witmer, Lawrence M. (1990). "Lesothosaurus, Pisanosaurus, and Technosaurus". In Weishampel, David B.; Dodson, Peter; Osmólska Halszka (eds.). The Dinosauria (1st ed.). Berkeley: University of California Press. pp. 416–425. ISBN 0-520-06727-4.
2. Dong, Z. (1989). On a small ornithopod (Gongbusaurus wucaiwanensis sp. now) from Kelamaili, Junggar Basin, Xinjiang, China. Vertebr. Palasiatica 27, 140-146. [In Chinese]
3. Sereno, P. C. (1991), Lesothosaurus, "fabrosaurids", and the early evolution of Ornithischia. J. Vertebr. Paleontol. 11, 168-197
4. Galton, P. M. (1978). Fabrosauridae, the basal family of ornithischain dinosaurs (Reptilia: Ornithopoda). Palaontol. Zeitschrift 52, 1-152
5. Gow, C. E. (1981). Taxonomy of the Fabrosauridae (Reptilia, Ornithischia) and the Lesothosaurus myth. South Africa J. Sci. 77, 43
6. Colbert, E. H. (1981). A primitive ornithischian dinosaur from the Kayenta Formation of Arizona. Museum Northern Arizona Bull. 53, 1-61